Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Cellular Respiration.

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Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Cellular Respiration

Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Energy Transformations

Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings E Transformations  Thermodynamics E flows into ecosystem as sunlight Light energy ECOSYSTEM CO 2 + H 2 O Photosynthesis in chloroplasts Cellular respiration in mitochondria Organic molecules + O 2 ATP powers most cellular work Heat energy Figure 9.2

Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Cellular respiration – Uses O 2 and sugar (glucose) – Yields ATP

Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings

mitochondria Mitochondrion Intermembrane space Outer membrane Free ribosomes in the mitochondrial matrix Mitochondrial DNA Inner membrane Cristae Matrix 100 µm

Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Overview of cellular respiration Glycolsis GlucosePyruvate ATP Substrate-level phosphorylation Mitochondrion

Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings ATP Substrate-level phosphorylation Mitochondrion Glycolsis GlucosePyruvate ATP Substrate-level phosphorylation Citric acid cycle

Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Electrons carried via NADH Glycolsis GlucosePyruvate ATP Substrate-level phosphorylation Electrons carried via NADH and FADH 2 Citric acid cycle Oxidative phosphorylation: electron transport and chemiosmosis ATP Substrate-level phosphorylation Oxidative phosphorylation Mitochondrion

Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Overview of cellular respiration Figure 9.6 Electrons carried via NADH Glycolsis Glucose Pyruvate ATP Substrate-level phosphorylation Electrons carried via NADH and FADH 2 Citric acid cycle Oxidative phosphorylation: electron transport and chemiosmosis ATP Substrate-level phosphorylation Oxidative phosphorylation Mitochondrion Cytosol

Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings 3 main processes in metabolism Electron shuttles span membrane CYTOSOL 2 NADH 2 FADH 2 2 NADH 6 NADH 2 FADH 2 2 NADH Glycolysis Glucose 2 Pyruvate 2 Acetyl CoA Citric acid cycle Oxidative phosphorylation: electron transport and chemiosmosis MITOCHONDRION by substrate-level phosphorylation by substrate-level phosphorylation by oxidative phosphorylation, depending on which shuttle transports electrons from NADH in cytosol Maximum per glucose: About 36 or 38 ATP + 2 ATP + about 32 or 34 ATP or Figure 9.16

Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings 2 possible pathways, with and without oxygen Glucose CYTOSOL Pyruvate No O 2 present Fermentation O 2 present Cellular respiration Ethanol or lactate Acetyl CoA MITOCHONDRION Citric acid cycle Figure 9.18

Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Evolutionary Significance of Glycolysis Glycolysis occurs in nearly all organisms Probably evolved before O 2 in the atmosphere