Protista: Applied Significances Diatoms  Diatoms are unicellular algae that have a unique glass-like wall  made of hydrated silica (silicon dioxide)

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Protista: Applied Significances Diatoms  Diatoms are unicellular algae that have a unique glass-like wall  made of hydrated silica (silicon dioxide) embedded in an or­ganic matrix.  The wall consists of two parts that overlap like a shoe box and its lid.  These walls provide effective protection from the crushing jaws of predators:

 In 2003, researchers in Germany found that live diatoms can withstand pressures as great as 1.4 million kglm2 equal to the pressure un­der each leg of a table supporting an elephant!  Much of the di­atoms' strength comes from the delicate lacework of holes and grooves in their walls; if the walls were smooth, it would take 60% less force to crush them.

A freshwater diatom (colorized SEM).

 With an estimated 100,000 living species, diatoms are a highly diverse group of protists.  They are a major component of plankton both in the ocean and in lakes: One bucket of water scooped from the surface of the sea may contain millions of these microscopic algae.  Like golden algae and brown algae, diatoms store energy in the form of a glucose polymer called laminarin. Some diatoms also store energy in oil molecules.

 Diatom populations may increase rapidly (bloom) when am­ple nutrients are available.  Although typically eaten by a variety of protists and invertebrates, during a bloom many diatoms escape this fate. When diatoms that are not eaten die, their bodies sink to the ocean floor.  This sinking process is part of a biological carbon pump that begins during photosynthesis when diatoms incorporate carbon into their bodies from car­bon dioxide in the air.

 Diatoms that sink to the ocean floor are not very likely to be broken down by bacteria and other decom­posers.  Hence, the carbon in their bodies is not returned to the air in the form of carbon dioxide respired by decomposers. In­ stead it is "pumped" to the ocean bottom.  With an eye toward reducing global warming by lowering atmospheric carbon diox­ide levels, some scientists advocate promoting diatom blooms by fertilizing the ocean with nutrients such as iron.  Other scientists question the wisdom of this strategy, noting that it is difficult to predict the effects of such large scale manipulations of ecological communities

Brown Algae  The largest and most complex algae are brown algae. All are multicellular, and most are marine.  Brown algae are especially common along temperate coasts, where the water is cool.  They are called brown algae due to their characteristic of brown or olive color to the carotenoids in their plastids.

 Many of the species commonly called ~seaweeds~ are brown algae. (Some large, multicellular species of red and green algae are also referred to as seaweeds.  Brown algae include species that have the most complex multicellular anatomy of all algae; some even have specialized tissues and organs that resemble those in plants.  But morphological and DNA evidence indi­cates that the similarities evolved independently in the algal and plant lineages and are thus analogous, not homologous.

Seaweeds: Adapted to life at the ocean's margins.  The sea palm {Pastelsial lives on rocks along the coast of the northwestern United States and western Canada.  The thallus of this brown alga is well adapted to maintaining a firm foothold despite the crashing surf.

 Brown algae are important commodities for humans. Some species are eaten, including Laminaria (Japanese "kombu"),Which is used in soups.  In addition, the gel-forming substance in the cell walls of brown algae, called algin, is used to thicken many processed foods, including pudding, ice cream, and salad dressing.

Red Algae

 Most red algae are multicellular.  Although none are as big as the giant brown kelps, the largest multicellular red algae are included in the informal designation "seaweed.  These are eaten one of these multicellular red algae, Porphyra  (Japanese "norn, as crispy sheets or as a wrap for sushi (Japanese food)

Green Algae  The grass-green chloroplasts of green algae have an ultra structure and pigment composition much like the chloroplasts of land plants.  Molecular systematics and cellular morphology leave little doubt that green algae and land plants are closely re­lated.  In fact, some systematists now advocate including green algae in an expanded ~plant" kingdom, Viridiplantae (from the Latin viridis, green).  Phylogenetically, this change makes sense, since otherwise the green algae are a para phyletic group.

Multicellular chlorophytes.

 Green algae are divided into two main groups, chlorophytes (from the Greek chloros, green) and charophytes.  More than 7,000 species of chlorophytes have been identified.  Most live in fresh water, but there are also many marine and some terrestrial species.