1.The Nitrogen Cycle 2.Amino Acid Biosynthesis 3.Urea Cycle 4.Amino Acid Catabolism 5.Molecules Derived from Amino Acid (from Lehninger P873- 882 )

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Presentation transcript:

1.The Nitrogen Cycle 2.Amino Acid Biosynthesis 3.Urea Cycle 4.Amino Acid Catabolism 5.Molecules Derived from Amino Acid (from Lehninger P )

The Nitrogen Cycle Page 810 nitrate nitrite nitrogenase Nitrite reductase Nitrate reductase

Nitrogen Fixation Is by Nitrogenase Complex Page 812

Three major reactions in all cells: 1.Carbamoyl-phosphate synthetase I NH 4 + +HCO ATP  carbamoyl-phosphate +2ADP+P i +2H + 2.Glutamate dehydrogenase 3.Glutamine synthetase The Fate of Ammonium Page 815

The Glutamate Dehydrogenase Reaction Page 815

The enzymatic reaction catalyzed by glutamine synthetase Page 815

Two principal pathways : 1. The GDH/GS pathway 2.GS/GOGAT in organisms confronting N limitation (GOGAT is glutamate synthase or glutamate:oxo- glutarate amino transferase) The Major Pathways of Ammonium Assimilation Lead to Glutamine Synthesis Page 816

The GDH/GS Pathway of Ammonium Assimilation Page 816 GDH: glutamate dehydrogenase GS: glutamine synthetase

The GS/GOGAT Pathway of Ammonium Assimilation Page 817

The Subunit Organization of Bacterial Glutamine Synthetase Page 818

Glutamine Synthetase (GS) in E. coli is regulated in three ways: 1.Feedback inhibition 2.Covalent modification (interconverts between inactive and active forms) 3.Regulation of gene expression Page 817,818 & 820

Allosteric Regulation of Glutamine Synthetase by Feedback Inhibition Page 819

Covalent Modification of GS: Adenylylation of Tyr397 in the Glutamine Synthetase Page 819

The Covalent Modification of GS Page 820

Gene Expression Regulates GS Page 820

Amino Acids Are Formed from  -keto Acids by Transamination Page 821

Human Dietary Requirements for Amino Acids Page 822

Amino Acid Biosynthesis Page 822

Chapter 25 Synthesis of Amino Acids in Bacteria

1 Page 822

Page 816 & 824

Urea cycle arginine Page 825

See page Page 822 & 831

Page 830

3 Pyruvate + glutamate  Alanine +  -KG Page 822 See P835 & P836

4 Page 822

Page 837

The Synthesis of S-adenosylmethionine (SAM) From Methionine Page 833

The Synthesis of S-adenosylmethionine (SAM) From Methionine Page 833

Page 822 5

Page 843 (also called Phenylalanine hydroxylase)

Some of the Aromatic Compounds Derived from Chorismate Page 841

Some of the Aromatic Compounds Derived from Chorismate Page 840

Page 842

Page 840 & 844 Histidine Biosynthesis and Purine Biosynthesis Are Connected by Common Intermediate

The Urea Cycle Page 827 Orithine transcarbamoylase Argininosuccinate synthetase Argininosuccinase arginase

Alanine Transports Ammonia from Muscles to the Liver

The TCA Cycle and Urea Cycle Can Be linked

Amino Acid Biosynthesis Inhibitors as Herbicides Page 838 *uncompetitive & noncompetitive inhibitor

Page 840 Overview of the Catabolism of Amino Acids in Mammals

Metabolic Degradation of the Common Amino Acids Page 841 & 845 Asn *glucogenic amino acid *ketogenic amino acid

Page 845

Page 848

Page 849 Maple syrup urine disease (楓漿尿病)

Page 849

Page 850

*Phenylketonuria(苯丙酮尿症) Page 843 & 848 Phenylalanine +  -KG transaminase Phenylpyruvate + glutamate (phenylalanine hydroxylase) Aspatame

*Alkaptonuria (黑尿症) Page 850 Homogentisate dioxygenase

*Maple syrup urine disease (楓漿尿病) (page 846) *Alkaptonuria (黑尿症)(page 848) *Phenylketonuria(苯丙酮尿症)(page 848)

1.Ammonotelic 2.Ureotelic 3.Uricotelic Page 850

Biomolecules Derived from Amino Acid (from Lehninger P ) Page 854 & Porphyrins

Page 855

Page 856 Aged erythrocyte Jaundice 黄疸

Page Creatine

3.Glutathione Page 858

4.Neurotransmitters Page 859 & 860

4.Polyamines Page 860 & 861

5.Nitric Oxide

phosphodiesterase GMP Viagra 威而鋼 Soluble Guanylyl Cyclases Are Receptors for Nitric Oxide Stripped away (sildenafil)