A Fish-Eating Enantiornithine Bird from the Early Cretaceous of China Provides Evidence of Modern Avian Digestive Features  Min Wang, Zhonghe Zhou, Corwin.

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A Fish-Eating Enantiornithine Bird from the Early Cretaceous of China Provides Evidence of Modern Avian Digestive Features  Min Wang, Zhonghe Zhou, Corwin Sullivan  Current Biology  Volume 26, Issue 9, Pages 1170-1176 (May 2016) DOI: 10.1016/j.cub.2016.02.055 Copyright © 2016 Elsevier Ltd Terms and Conditions

Figure 1 Enantiornithine Bird, IVPP V22582, Preserving a Pellet Containing Fish Bones Abbreviations are as follows: bc, braincase; fr, frontal; fu, furcula; gp, gastric pellet; lc, left carpometacarpus; lco, left coracoid; lf, left femur; lh, left humerus; lp, left pubis; lr, left radius; ls, left ischium; lt, left tibiotarsus; lu, left ulna; rc, right carpometacarpus; rco, right coracoid; rf, right femur; rfi, right fibula; rh, right humerus; ri, right ilium; rp, right pubis; rr, right radius; rs, right ischium; rt, right tibiotarsus; rta, right tarsometatarsus; ru, right ulna; ps, premaxillary symphysis; st, sternum; sy, synsacrum. Scale bar, 10 mm. See also Figure S1 for interpretative drawing. Current Biology 2016 26, 1170-1176DOI: (10.1016/j.cub.2016.02.055) Copyright © 2016 Elsevier Ltd Terms and Conditions

Figure 2 Pellets Containing Fish Bones Preserved in Birds from the Early Cretaceous, ∼120 Million Years Old, Jehol Biota (A and B) Enantiornithine bird skeleton (IVPP V22582) with a preserved pellet: view of the pellet overlain by the right humerus (A) and close-up of the fish bones (B). (C and D) Two pellets preserved in the ornithuromorph bird Piscivoravis lii (IVPP V17078): one close to the dentary (C) and the second in the abdominal region (D). fb, fish-bone pellet; hu, humerus. Scale bars, 10 mm (A, C, and D) and 1 mm (B). Current Biology 2016 26, 1170-1176DOI: (10.1016/j.cub.2016.02.055) Copyright © 2016 Elsevier Ltd Terms and Conditions

Figure 3 A Sketch Reconstruction of the Fish-Eating Enantiornithine Bird Represented by IVPP V22582 The stomach is two chambered, with a distinct proventriculus and gizzard, and the indigestible fish bones are being ejected as a pellet; the presence of a crop is likely but uncertain in enantiornithines. The gizzard is depicted in anterior view (cross-section) to show the narrow pyloric opening into the small intestine. cr, crop; es, esophagus; gz, gizzard; pe, pellet; pr, proventriculus. Current Biology 2016 26, 1170-1176DOI: (10.1016/j.cub.2016.02.055) Copyright © 2016 Elsevier Ltd Terms and Conditions