Mean heart rate (fh, A), rate of oxygen of consumption (V̇O2, B) and mass-specific V̇O2 (sV̇O2, C) of 22 king penguins walking at different speeds on a.

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Mean heart rate (fh, A), rate of oxygen of consumption (V̇O2, B) and mass-specific V̇O2 (sV̇O2, C) of 22 king penguins walking at different speeds on a treadmill. Mean heart rate (fh, A), rate of oxygen of consumption (V̇O2, B) and mass-specific V̇O2 (sV̇O2, C) of 22 king penguins walking at different speeds on a treadmill. The birds are in four groups, females (squares, N=5, FD) and males (circles, N=5, MD) departing from the colony and females (triangles, N=6, FA) and males (crosses, N=6, MA) arriving at the colony after a foraging shift. The vertical and horizontal lines at each point represent ± s.e.m. G. Froget et al. J Exp Biol 2001;204:2133-2144 © The Company of Biologists Limited 2001