Marginal means (±s.e.m.) of temperatures for four tissues in 13 king penguins while fasting in water. Marginal means (±s.e.m.) of temperatures for four.

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Marginal means (±s.e.m.) of temperatures for four tissues in 13 king penguins while fasting in water. Marginal means (±s.e.m.) of temperatures for four tissues in 13 king penguins while fasting in water. Values for the flank (A), the back (B), the brood patch (C) and the abdominal cavity (D) are shown, which were computed from separate LMMs for daytime (light colour) and nighttime periods (dark colour; Table 1, see text). Values that do not share the same letter are significantly different from each other (post hoc Tukey’s HSD test; P<0.05). n.s., not significant. Agnès Lewden et al. J Exp Biol 2017;220:4600-4611 © 2017. Published by The Company of Biologists Ltd