SUPPORT AND TRANSPORT IN PLANTS ANATOMY OF DICOT PLANTS

ADVENTITIOUS ROOT SYSTEM The Root Two types of root systems - tap root system - adventitious root system TAP ROOT SYSTEM ADVENTITIOUS ROOT SYSTEM - Arises from the radicle of the embryo - Arises from any part of the plant except from the radicle of the embryo - Main root can be distinguished - Main root cannot be distinguished

Functions of the roots They anchor the plant firmly to the ground. They absorb water and mineral salts.

External Structure of Root Tip Root cap is made up of parenchyma cells. It protects the inner layers of the root. The growing point is made up of meristematic cells. Some of the new cells produced replace the parenchyma cells of the root cap which were damaged. Region of elongation – the cells comes from the growing point which begin to elongate. The root hair region has root hairs, which absorb water and mineral salts. Internally the cells in this region begin to differentiate into epidermis, parenchyma, sclerenchyma, xylem and phloem Mature region has side roots and fully differentiated tissues.

Transverse section of young dicot root, in plan

Transverse section of a young dicot root in detail

Enlarged Root Hair

Functions of the various root tissues . The root hairs of the epidermis absorb water and mineral salts. The parenchyma of the cortex stores starch and the intercellular spaces allow water and mineral salts to pass through The passage cells of the endodermis direct water into the xylem of the stele. The pericycle gives rise to side roots The xylem transports water and mineral salts from the roots to the stem and leaves. It also provides strength to the plant. The phloem transports manufactured food from the leaves to the roots. The vascular cambium gives rise to additional xylem and phloem as the root grows in thickness

The Stem Functions of the stem: They hold the leaves in a favourable position to receive light for photosynthesis. They hold the flowers in a favourable position for pollination. Most stems contain chlorophyll and they therefore manufacture food by the process of photosynthesis. They store food and water They transport water and mineral salts from roots to leaves They transport food from the leaves to the roots

External structure of a dicot stem

Transverse section of a young dicot stem in plan

Transverse section of a young dicot stem in detail

Functions of the various stem tissues The cuticle reduces water loss through transpiration The cuticle is transparent allowing light to pass through for photosynthesis. The epidermis may have guard cells to allow gaseous exchange while at the same time reducing water loss through transpiration. The hypodermis of sclerenchyma or collenchyma strengthens the stem and helps keep it upright. The parenchyma of the cortex stores food. The intercellular air spaces between the parenchyma cells allow for transport of water and gases. The endodermal cells stores starch. The sclerenchyma fibres gives strength to the plant and help to keep it upright. Xylem transports water and mineral salts from the roots to the leaves Phloem transports manufactured food from the leaf to the roots

Secondary growth of the stem Increase in thickness of the stem is called secondary growth or secondary thickness. Secondary thickness takes place in 3 steps: Formation of secondary xylem and secondary phloem Formation of cork Formation of lenticels

Formation of secondary xylem and secondary phloem Parenchyma cells between the vascular bundles begin to divide. This with vascular cambium forms a complete ring of cambium Entire cambium divides to form secondary xylem on the inside and secondary phloem on the outside. Every year a new ring of secondary xylem is formed. These are called the annual rings. (can tell the age of the tree by counting annual rings) The xylem cells making up the older annual rings becomes clogged and are no longer able to transport water and mineral salts. They become heartwood.

Transverse section of stem after a complete ring of cambium has been formed

Formation of Cork Certain parenchyma cells in the cortex begin to divide and form cork cambium The cork cambium divides to form cork cells on its outside. The cork cells have suberin laid down on their cell walls.

Annual Rings

Formation of Lenticles In woody stems the stomata are blocked by the presence of cork cells The epidermis of woody stems breaks up to form tiny pores called lenticles which allow gaseous exchange.

XYLEM Xylem tissue is made up of four types of cells - xylem vessels - xylem tracheids - xylem sclerenchyma - xylem parenchyma Xylem vessels and xylem tracheids are highly specialised cells. Xylem vessels are dead cells. They are long, cylindrical cells. Their cross walls are perforated or completely absent and form continuous tubes from roots to leaves. Xylem vessels have thick walls made up of lignin. Lignin is laid down in various patterns Annular thickening ( lignin laid in circles) Spiral thickening ( lignin laid in spiral) Scalariform thickening (laid in ladder form) Pitted thickening (laid in pits) Xylem tracheids are similar to xylem vessels except that the ends are tapered and cross walls are always present

Xylem Vessels Xylem Tracheids Pitted Thickening Scalariform Thickening Annular thickening Spiral Thickening Xylem Tracheids

Adaptation of xylem to transport Cross walls are perforated or completely absent – this forms xylem roots forms continuous tubes with the xylem stems and leaves. Xylem vessles have no living contents – allows the water to flow freely inside. The walls of the xylem vessels and tracheids are not completely thickened – the unthickened portions and bordered pits allow water to move across the root and stem Adaptation of xylem to provide strength the vessels and tracheids are thick-walled – they have a secondary wall of lignin the vessels and tracheids are round in cross-section – round structures give additional strength

Phloem Phloem tissue is made up of four types of cells: Sieve tubes Companion cells Phloem parenchyma Phloem sclerenchyma Sieve tubes and companion cells are highly specialised cells Sieve tubes are living cells with long cylindrical cells. Their cross walls are perforated to form sieve plates They are thin walled cells They contain strands of cytoplasm which run through the sieve plates from one cell to the next Companion cells lie next to the sieve tubes – they are thin walled, with cross walls and has a well-defined nucleus

Phloem tissue

The function of phloem tissue The sieve tubes transport manufactured food from the leaves to the all parts of the plant. The companion cells probably control the activities of the sieve tubes

Collenchyma and Sclerenchyma

Absorption and Transport of water and mineral salts by plants Water enters the root hairs of the epidermis of roots by osmosis POINTS TO REMEMBER THE SOIL WATER IS THE DILUTE SOLUTION THE CELL SAP HAS CONCENTRATION SOLUTION BECAUSE OF HIGH CONCENTRATION OF SALTS. THE CELL MEMBRANE AND TONOPLAST IS SEMI-PERMEABLE MEMBRANE OSMOSIS: MOVEMENT OF WATER MOLECULES FROM HIGH WATER POTENTIAL TO LOW WATER POTENTIAL THROUGH SEMI PERMEABLE MEMBRANE.

Lateral transport of water and mineral salts to the stele of the root

Upward movement of water and mineral salts through the stem from Roots to Leaves

Forces responsible for the upward movement of water Transpirational pull Root pressure Capillarity

Forces responsible for the upward movement of water Transpirational Pull: Water evaporates from leaves Creates diffusion pressure gradient between outside air and xylem Tension exists in the water column extending from the roots to the leaves Adhesive and cohesive forces keep the column continuous Thus water is continually being pulled upward as water evaporates from the leaves Upward pull = transpirational pull

Forces responsible for the upward movement of water Root pressure Cell sap of root hair always has low water potential when compared to soil water Water enters root hair by osmosis Continuous osmosis causes pressure forcing water a short distance up the stem Capillarity Spontaneous movement of water up narrow tubes because of adhesion and cohesion is called capillarity Lumen of xylem vessels and tracheids are very narrow therefore capillarity takes place Upward distance is very short

Structural modifications of leaves to reduce water loss (Internal factors limiting transpiration) Sunken stomata: In some plants stomata may be sunken or small. Thickened cuticle: A thick cuticle on the surface of leaves reduces the rate of transpiration. Hair on leaves: the sheen r shine of the hairs of some leaves reflect sunlight and reduces transpiration. Hairs also trap water vapour, restricting water loss. Shape, size and arrangement of leaves: The shape and size of leaves determine the total surface area exposed to the environment, and hence influence the rate of transpiration. The leaves of some plants are arranged in such manner that they shade each other or overlap each other. In this water loss is restricted

Environmental factors affecting the rate of transpiration Humidity: High humidity decreases transpiration rate Wind: Increases evaporation and thus transpiration increases. Wind removes the water vapour around leaf thus increasing water vapour pressure gradient. Temperature: Increases evaporation and thus transpiration Light: generally causes an increase in the rate of transpiration

Wilting: Loss of turgidity of the cells results in leaves and Sometimes the stems, becoming limp, causing them to droop. Then the plant is called wilting. Guttation: Loss of liquid water through tiny pores, called hydathodes on margin of leaf. Guttation Wilting Plant

Economic uses of plants related to their anatomy: Paper Clothing Timber for furniture-making and building construction Basket-making Thatching, matting and broom-making

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