Competition
Competition is an interaction between individuals, brought about by a shared requirement for a resource in a limited supply, and leading to a reduction in the survivorship, growth, and/or reproduction of the competing individuals
Competition
Two key points: 1.Resource being competed for must be limited in supply 2. Interaction is always negative
Competition Intraspecific competition - competition between members of the same species - thought to be especially strong because members of the same species have the same resource requirements – regulates population size Interspecific competition - competition between members of different species - can be very strong as well – also regulates population size, structures communities, influences natural selection
Mechanisms of Competition 1) exploitation competition (aka resource or scramble) - individuals exert negative influence on each other by using up resources that each need - competition is mediated indirectly through resource depletion - individuals don't need to come into contact to compete 2) interference competition (aka contest competition) - competitors directly interact with each other in the course of seeking a common resource
Coral reef community – Great Barrier Reef
Competitive Asymmetries Although competition has negative effects on the population as a whole, some individuals are more negatively effected than others
Flax
Orchard Grass
Territoriality Territoriality occurs when there is an active interference such that a more or less exclusive area, the territory, is defended against intruders by a recognizable pattern of behavior - individuals or groups are spaced out more than would be expected from a random occupation of suitable habitats
Golden-winged sunbird
Limpet – Patella cochlear
Patella grazing on crustose algae (A)
Corn
Law of Constant Final Yield
Georgii F. Gause
Paramecium
Competitive Exclusion Principle "Two competing species cannot coexist in a stable environment if both require the same limiting resource." - if two competing species do coexist in a stable environment, they do so because of niche differentiation - their realized niches are slightly different and thus competition between them is lessened
Robert MacArthur,
Caveats to Resource Utilization Curves 1) Are we examining right resource? Is something else really limiting? 2) Is there variation in when the resource is limiting? If it is only limiting occasionally, then it may not matter much. 3) How do we quantify resource use by species in nature? Do resource utilization curves have to be normal? Does competition change during life history changes in the species? 4) How does overlap really affect the species? Is overlap really only on 1 dimension?