The Human Visual System Vision Class The main topics covered here were: photoreceptors and ganglion cells in the retina. The photoreceptors are the input, the ganglion are the output to the brain. We also discussed the role of the ganglion cells in edge detection.
The Human Brain: Overall View Visual information goes from the eyes via the LGN in the thalamus to the back of the cortex
Each half of the visual field goes to a different hemisphere of the brain
Visual pathway
Hemifield neglect
Eye
Recording
Recording from a Neuron
Neurons Purkinje cells from cerebellum, dendrites showing calcium concentration
Synaptic connection
Synapse
Receptors Density - Fovea
Photoreceptors
Photo response Retinal (the chromophore) + opsin (rodopsin, cone-opsin) 11-cis retinal + photon → all trans retinal Activates transducin (a G- protein) Activates phosphodiesterase Breaks cGMP cGMP keep ionic channels open, now some close (one retinal → 500 cGMP) Cell hyperpolarization
Image Capture Huge dynamic range – μW/cm 2 Photons: poisson process. Noisy at low levels For low light: large receptors, slow integration Rods/cones, local adaptation,change of amplitude and time constant, motion deblur
Dynamic Range
Visual receptor types
Retina Mosaic
Color Mixing
Edge Detection by Z.C. Peak in f’(x) z.c. in f’’(x) Noise: Asin(wx) > -w 2 cos(wx) Convolve with a Gaussian (optimal) d 2 /dx 2 (G*f) = d 2 /dx 2 (G) * f In 2-D: d 2 /dx 2 + d 2 /dy 2
Edge Detection by Z.C.
Z.C. at Two Scales
Wheel Edges 4 Resolutions
END