Phloem Translocation and Assimilate Partitioning

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Presentation transcript:

Phloem Translocation and Assimilate Partitioning HORT 301 – Plant Physiology November 2, 2009 Taiz and Zeiger, Chapter 10 paul.m.hasegawa.1@purdue.edu Assimilate translocation in the phloem from source-to-sink is bidirectional

Phloem sap composition

Phloem tissue is adjacent to the xylem Stem (perennial) Leaf

Sieve elements and companion cell

Companion cell types Ordinary companion cell Transfer cell Intermediary cell

Source to sink translocation Proximity Vascular interconnection Sink strength

Pressure flow model for phloem translocation from source to sink Source – phloem loading increases ψp Sink – phloem unloading decreases ψp Ψp gradient between source and sink drives solution movement

Phloem loading from source cells and unloading into sink cells Photo-assimilate movement Symplast (active) and apoplastic (active) phloem loading

Apoplastic (active) phloem loading Summary of apoplastic and symplastic phloem loading

Phloem unloading and storage in sinks

Photosynthetic product allocation and partitioning Allocation – regulation of carbon assimilate distribution into various metabolic pathways or sink cells Partitioning – distribution of carbon assimilates within plants

Translocation from source to sink