Cnidarian Diversity

Phylum Cnidaria Class Anthozoa Class Hydrozoa Class Scyphozoa Class Cubozoa Corals Anemones Hydra Portuguese Man-Of-War Stinging Limu Fire Coral True jellyfishBox jellies Sea wasps

Class Anthozoa Subclass Hexacorallia Order Antipatharia Subclass Alyconaria Order Zoanthidae Order Actinaria Order Scleractinia ZoanthidsSea anemonesStony coralsPrecious black coral Wire coral Sea fans Sea whips Sea pens Organ pipe coral Precious gold coral

General Characteristics of Cnidaria Stinging cells Two stages in life cycle (polyp & medusa) Blind sack gut Radial symmetry Diploblastic Hydrostatic skeleton Nerve net Many colonial, some solitary forms

polyps and medusa stage, although polyp stage is dominant. gut cavity of polyp is simple, lacking a pharynx and not divided by mesenteries. tetramerous (four-part) radial symmetry. gonads are ectodermal (found in the epidermis). medusa stage may possess specialized balance organs called statocysts and photosensitive organs called ocelli. solitary or colonials; some colonial forms highly polymorphic. includes hydroids (“stinging limu”), fire coral, pink coral, and siphonophores. Class Hydrozoa (Gr. hydra, water serpent)

The Cnidarian Life Cycle The Hydrozoan Life Cycle

Hydrozoan Colony

Hydrozoan Colonies “Stinging Limu”

Hydrozoan Medusa

Hydrocorals

Fire Coral

By-the-Wind-Sailor A Floating Colony of Polyps

Coloniality Man-of-War Phylum Cnidaria

Close Up of a Portuguese Man-Of-War

life cycle with both polyps and medusae, but medusae dominate with polyp stage reduced or absent. polyp stage (scyphistoma) goes through strobilization to produce young medusa. bell margin lacks a velum. tetramerous (= four-part) radial symmetry. gut divided into a complex system of radial canals. some with a simple single mouth, but many with thousands of microscopic “mouths” at the ends of oral arms. gonads endodermal (found in the gastrodermis). specialized sense organs called rhopalia with ocelli & statocysts. includes some 200 marine species. "true" sea jellies. Class Scyphozoa (Gr. skyphos, cup)

planula scyphistoma strobila ephyra adult medusa gametes Scyphozoan Life Cycle strobila scyphistoma

Sea Jellies

Moon Jelly Anatomy

polyps and medusae stages, but medusae dominate with polyp stage reduced. polyp stage develops directly into medusa. bell margin with a velarium. tetramerous (= four-part) radial symmetry; bell cube- shaped with tentacles arising from each corner. gonads endodermal (found in the gastrodermis). specialized sense organs called rhopalia with ocelli & statocysts. includes some 15 marine species. includes box jellies and sea wasps. Class Cubozoa (Gr. kybos, a cube)

Box Jellies Seawasp

Box Jelly Anatomy

lack medusa stage entirely (polyp forms only) mouth with a tubular pharynx that projects inward into the gut large gut cavity divided by mesenteries that radiate inwards from the body wall gonads endodermal, borne on the mesenteries hexamerous (6-part) or octamerous (8-part) radial symmetry or biradial (modified radial symmetry that limits the number of planes that can divide the body into equal halves includes sea anemones, “true” stony corals, sea fans, sea pens, organ pipe coral, precious black coral, & zoanthids (Gr. anthos, flower) Class Anthozoa

Subclass Zoantharia Order Actinaria Sea Anemones

Sea Anemone Anatomy

Subclass Hexacorallia Order Zoanthidae No hard skeleton May be tough and leathery Shallow water forms Encrusting forms Some with zooxanthellae Polyp stage only

Subclass Hexacorallia Order Antipatheria Black Coral & Wire Coral Wire coral Black coral

Subclass Hexacorallia Order Scleractinia “True” Stony Corals Hermatypic- reef building, zooxanthellae Common species: Acroporidae- table Acroporidea- rice Agariciidae- flat lobe, corrugated Faviidae- crust, ocellated Fungiidae- humpback, mushroom Pocilloporidae- lace, antler, cauliflower Poritidae- finger, lobe, plate

“True” Stony Corals lobe finger mushroom Porites rus

8 branched tentacles surrounding the mouth Both hard and soft forms exist Ahermatypic- non reef building, no photosynthesis Five Orders found in Hawaii: 1.Stolonifera 2.Pennatulacea (sea pens) 3.Alcyonacea (soft corals) 4.Telestacae (snowflake corals) 5.Gorgonacea (sea fans) Subclass Alyconaria Octocorals

Sea pen

Organ pipe coral