Consumption and Secondary Production Reading: pp. 411-416 (5 th ) A. Food chains and food webs Grazing Detrital (decomposer) B. Energy budget - flow of.

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Consumption and Secondary Production Reading: pp (5 th ) A. Food chains and food webs Grazing Detrital (decomposer) B. Energy budget - flow of energy through an ecosystem C. Trophic levels and ecological pyramids D. Efficiency of energy transfer Consumption, Assimilation, Growth, Secondary production E. How can we determine food web relationships?

Why are big, fierce animals so scarce? Where does the energy come from that fuels ecosystems? What is the fate of that energy? How does it affect the distribution and abundance of organisms of different types?

A. Food chains are simplifications of food webs Simplified food web

B. Energy Budget: Source and fate of energy

ECOSYSTEM PRINCIPLES 1- Energy flow 2- Chemical cycling ENERGY- Cannot be recycled, flows through ecosystems, from an external source, enters as light exits as heat. MATTER- Cycles within ecosystems. Autotrophs Heterotrophs Energy flow and chemical cycling described by grouping species in a community into trophic levels according to main source of nutrition and energy Tertiary consumers, etc. PHOTO, chemo (decomposers) C&R 54.1

B. Energy Budget: Source and fate of energy Points: 1. GPP > NPP > NEP 2. Energy flow is one-way - once used, it is dissipated as heat

C-cycle: the somewhat more detailed version

C. Trophic pyramids Classic food chain 2. Rule of thumb: 10% energy transfer between trophic levels 1. Trophic levels: Primary producers, herbivores, carnivores (predators), omnivores, detritivores

Consequences for diversity

Can this ever happen with pyramids based on energy flow (productivity)? 1 o producers herbivores carnivores Biomass at each trophic level Inverted trophic pyramids

D. Efficiencies of energy transfer Why is biomass of animals so small? Where does all the energy go? Why is transfer efficiency so low?

C-cycle: the somewhat more detailed version

Availability of energy for growth ConsumedUnconsumed 1 o Prod Production Respiration Assim. Assimilated Urine Consumed Feces So, P = C - R - F - U

Trophic energy losses: a Michigan old-field

Availability of energy for growth: Depends on efficiency of transfer ConsumedUnconsumed 1 o Prod Production Respiration Assim. Assimilated Urine Consumed Feces Assimilation efficiency = A/C Production efficiency = P/A Growth efficiency = P/C = P/A * A/C

Assimilation, production, and growth efficiencies for homeotherms and poikilotherms Smith (1998) Table 11.3, p. 181

Basal metabolic rate – endos & ectos

E. How can we determine food web relationships? Observation Gut/feces content You are what you eat, isotopically speaking.

Fig – Human consumption of corn

Fig

Fig – food sources of the ribbed mussel.