Course web site www.clas.ufl.edu/users/mcmack/2008.

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Course web site

Module 1: Evolution MonthDayTopic Aug28History of life on Earth 30Timeline construction Sept1Theory of evolution 4Holiday 6Evidence for evolution 8Mechanisms of evolution I 11Mechanisms of evolution II 13Speciation

Outline 1.What is life? 2.How might it have arisen? 3.What has happened since then?

Time Cosmological Geological Evolutionary Ecological Physiological

4.5 billion years ago, the earth was born

Life is...what?

Life shares Chemical composition – CHON(SP) Carbon Hydrogen Oxygen Nitrogen Sulfur Phosphorus Iron, Zinc, Calcium, Sodium...

Life shares Biochemistry in liquid water –Proteins –Lipids –Carbohydrates –Nucleic acids

Life shares Use of matter and energy –Energy stored and released by breaking chemical bonds –Autotrophs –Heterotrophs –Metabolism –Enzymes –Homeostasis

Life shares Cells –Membranes –Cytoplasm –Organelles –Nucleus A prokaryote cell...

Life shares Reproduction –Heredity –Variation

Life shares Ecology!!!!!

Life shares... 1.Chemical composition-CHON(SP) 2.Biochemistry 3.Use of matter and energy 4.Cells 5.Reproduction 6.Ecology

How did life start?

Causal mechanism

The biogenesis paradox Life arises from life... But what about the first organisms?

How did life start? In a very different earth environment –Reducing atmosphere—No O 2 ! –Lots of H 2 O –Lightening, volcanoes, ultraviolet rays

The primordial soup hypothesis 1.Abiotic synthesis of simple organic molecules

The primordial soup hypothesis 1.Abiotic synthesis of simple organic molecules 2....complex organic molecules

The primordial soup hypothesis 1.Abiotic synthesis of simple organic molecules 2....complex organic molecules 3.Origin of self-replicating molecules

The primordial soup hypothesis 1.Abiotic synthesis of simple organic molecules 2....complex organic molecules 3.Origin of self-replicating molecules 4.Formation of pre-cells

The big leap From pre-cells to life... Lots of time for natural selection

Life first appears in the fossil record about 3.8 billion years ago

Changes on Earth that have shaped life… Atmosphere (O 2, UV) Continental drift Climate Volcanoes Extraterrestrial bombardment

History has been punctuated by mass extinctions

1 st mass extinction 440 mya: ~75% of species disappear Sudden global cooling? Predatory arthropod

2 nd mass extinction 370 mya: ~75% of species disappear Continental collision or global climate change?

3 rd mass extinction 245 mya: ~96% of species disappear The greatest event of all time! Extraterrestrial impact?

4 th mass extinction 210 mya: ~65% of species 5 th mass extinction 65 mya: ~76% of species Extraterrestrial impact or volcanism?

The 6 th mass extinction... Last 10,000 years –30% of earth’s species gone by year 2100