Lecture 5 Outline (Ch. 35) Overview - Monocots vs. Dicots II. Plant tissues A. Dermal B. Vascular C. Ground Cell Types: Parenchyma, Collenchyma, Sclerenchyma III. Plant organs A. Roots B. Stems C. Leaves IV. Plant Growth A. Meristems B. Primary vs. secondary i. Leaves ii. Stems iii. Roots V. Preparation for next lecture

Plant Classification – Monocots vs. Dicots Basic categories of plants based on structure and function

Three Basic Plant Organs: Roots, Stems, and Leaves Plant “bodies” Plants, like multicellular animals, have organs composed of different tissues, which in turn are composed of cells Shoot system Leaf Stem Three Basic Plant Organs: Roots, Stems, and Leaves Root system

Setting the scene - animal bodies Organs Tissues Cells

Plant Tissues Dermal tissue Ground Vascular Each plant organ has dermal, vascular, and ground tissues Each of these three categories forms a tissue system

Plant Tissues 1) Dermal Tissue System Outer covering Protection 2) Vascular Tissue System “Vessels” throughout plant Transport materials 3) Ground Tissue System “Body” of plant Photosynthesis; storage; support

What type of tissue transports fluids in plants? A. Dermal B. Roots C. Vascular D. Stems E. Ground

Plant Tissues - Dermis Dermal Tissue System (Outer Covering of Plant): 1) Epidermal Tissue (epidermis): Outermost layer Cuticle: Waxy covering Reduces evaporation/predation Root Hairs: extended root surface Increase absorption 2) Peridermal Tissue (periderm): Only in woody plants (“bark = dead cells”) Protection; support

Plant Tissues - Dermis Special Dermal Cells – Trichomes & Root hairs Hairlike outgrowths of epidermis Keep leaf surfaces cool and reduce evaporation Roots hairs Tube extensions from epidermal cells Greatly increase the root’s surface area for absorption

Plant Tissues - Dermis Special Dermal Cells – Guard Cells Stomata Stomata Guard cells Guard cells Epidermal cell Epidermal cell a. c. 4 µm 200 µm Paired sausage-shaped cells Flank a stoma – epidermal opening Passageway for oxygen, carbon dioxide, and water vapor Stoma Stoma Epidermal cell Epidermal cell Guard cells Guard cells b. 71 µm

Plant Tissues - Vascular 1) Xylem (dead at maturity) cells: Tracheids: Narrow, tube-like cells Vessel Elements: Wide, tube-like cells

Plant Tissues - Vascular Vascular Transport System 1) Xylem (dead at maturity): - Moves water & minerals from roots to shoots

Plant Tissues - Vascular Sieve Tubes: Wide, tube-like cells B) Companion Cells: support and regulate sieve tubes 2) Phloem (living at maturity) cells:

Plant Tissues - Vascular Vascular Transport System 2) Phloem (living at maturity) - Moves water, sugar, amino acids & hormones

Vasculature - Comparisons Monocots and dicots differ in the arrangement of vessels in the roots and stems Dicots Monocots Stem Root

Plant Tissues – Ground Tissue Tissues that are neither dermal nor vascular are ground tissue Ground tissue internal to the vascular tissue is pith; ground tissue external to the vascular tissue is cortex Ground tissue includes cells specialized for storage, photosynthesis, and support Some major types of ground cells: Parenchyma Collenchyma Sclerenchyma

Plant Tissues - Ground Ground Tissue System (“Body” of Plant): 1) Parenchyma (most abundant): Thin-walled cells; living plant metabolism: Photosynthesis; hormone secretion; sugar storage FIGURE 42-5 The structure of ground tissue (a) Parenchyma cells are living and serve many functions. They have thin, flexible primary cell walls. These parenchyma cells are used for starch storage in a potato. (b) Collenchyma cells are living and have thickened, but somewhat flexible, primary walls. They help support the plant body (as seen in this celery stalk). (c) Sclerenchyma cells have thick, rigid secondary cell walls and die after they differentiate. Illustrated are "stone cells" that give pear fruit its slightly gritty texture. Parenchyma cells in Elodea leaf,(w/chloroplasts)

Plant Tissues - Ground Ground Tissue System (“Body” of Plant): 2) Collenchyma: Thick-walled (uneven); living Offers support (flexible & strong) FIGURE 42-5 The structure of ground tissue (a) Parenchyma cells are living and serve many functions. They have thin, flexible primary cell walls. These parenchyma cells are used for starch storage in a potato. (b) Collenchyma cells are living and have thickened, but somewhat flexible, primary walls. They help support the plant body (as seen in this celery stalk). (c) Sclerenchyma cells have thick, rigid secondary cell walls and die after they differentiate. Illustrated are "stone cells" that give pear fruit its slightly gritty texture. Collenchyma cells sunflower

Plant Tissues - Ground Ground Tissue System (“Body” of Plant): Sclereid cells in pear (LM) Fiber cells in ash tree Cell wall 3) Sclerenchyma: Thick, hard-walled; Dead Offer support (e.g. hemp fibers; nut shells) FIGURE 42-5 The structure of ground tissue (a) Parenchyma cells are living and serve many functions. They have thin, flexible primary cell walls. These parenchyma cells are used for starch storage in a potato. (b) Collenchyma cells are living and have thickened, but somewhat flexible, primary walls. They help support the plant body (as seen in this celery stalk). (c) Sclerenchyma cells have thick, rigid secondary cell walls and die after they differentiate. Illustrated are "stone cells" that give pear fruit its slightly gritty texture.

Which is a plant cell type? A. Ground B. Pith C. Sieve-tubes D. Cortex E. Stem

Self-Check Cell types Tissue Types Organ Types Parenchyma Ground Stems Collenchyma Vascular Shoots (& flowers) Sclerenchyma Dermal Roots Tracheids Vessels elements Sieve-tubes Companion cells Epidermal Guard cells

Roots - Overview Roots need sugars from photosynthesis; Shoots rely on water and minerals absorbed by the root system Root Roles: - Anchoring the plant - Absorbing minerals and water - Storing organic nutrients

Roots - Comparisons Taproots: Fibrous roots: Typical of dicots, primary root forms and small branch roots grow from it In monocots mostly, primary root dies, replaced by new roots from stem

Roots – Structure and Development Four regions: Root cap Protection, gravity detection Zone of cell division Mitotic divisions Zone of elongation Cells lengthen, no division Zone of maturation Cells differentiate, outer layer becomes dermis

Roots – Structure and Development In maturation zone, Casparian strip forms – waterproof barrier material surrounding vasculature

Roots – Structure and Development Epidermis Primary phloem Primary xylem Pith Monocot Eudicot Endodermis Cortex Pericycle 48 µm 385 µm 8 µm Location of Casparian strip

Roots – Many Plants Have Modified Roots Prop roots “Strangling” aerial roots Storage roots Buttress roots Pneumatophores Roots – Many Plants Have Modified Roots Figure 35.4 Modified roots Water storage

Stems - Overview Stem: an organ made of Apical bud Node Internode Apical bud Shoot system Vegetative shoot Axillary Stem Stem: an organ made of An alternating system of nodes, points at which leaves attach Internodes, stem length between nodes Axillary bud - structure that can form a lateral shoot, or branch Apical/terminal bud - located near the shoot tip, lengthens a shoot Apical dominance maintains dormancy in most nonapical buds

Vasculature - Stems In most monocot stems, the vascular bundles are scattered throughout the ground tissue, rather than forming a ring Phloem Xylem Sclerenchyma (fiber cells) Ground tissue connecting pith to cortex Pith Cortex 1 mm Epidermis Vascular bundle Cross section of stem with vascular bundles forming a ring (typical of eudicots) (a) Key to labels Dermal Ground Cross section of stem with scattered vascular bundles (typical of monocots) (b) bundles tissue Figure 35.17 Organization of primary tissues in young stems Dicot Monocot

Stems – Structure and Development Stems have all three types of plant tissue Grow by division at meristems Develop into leaves, other shoots, and even flowers Leaves may be arranged in one of three ways

Stems – Many Plants Have Modified Stems Rhizomes Bulbs Storage leaves Stem Stolons Stolon Tubers Figure 35.5 Modified stems

Leaves - Overview Shoot system Leaf Blade Petiole The leaf is the main photosynthetic organ of most vascular plants Leaves generally have a flattened blade and a stalk called the petiole, which joins the leaf to a node of the stem

Leaves – Structure and Development Leaves are several layers thick – each with different cell types

Leaves – Structure and Development Most dicots have 2 types of mesophyll Palisade mesophyll high photosynthesis Spongy mesophyll air spaces for gas & water exchange Monocot leaves have 1 type of mesophyll

Leaves Leaf epidermis contains stomata - allow CO2 exchange Stomata flanked by two guard cells, control open vs. closed Key to labels Dermal Ground Vascular Cuticle Sclerenchyma fibers Stoma Bundle- sheath cell Xylem Phloem (a) Cutaway drawing of leaf tissues Guard cells Vein Lower epidermis Spongy mesophyll Palisade Upper Stomatal pore Surface view of a spiderwort (Tradescantia) leaf (LM) Epidermal (b) 50 µm 100 µm Air spaces Guard cells Cross section of a lilac (Syringa)) leaf (LM) (c) Figure 35.17 Organization of primary tissues in young stems

Leaves - Comparisons Monocots and dicots differ in the arrangement of veins, the vascular tissue of leaves Most dicots have branch-like veins and palmate leaf shape Monocots have parallel leaf veins and longer, slender blades

Leaves – Plants have modified leaves for various functions Tendrils Spines Storage leaves Figure 35.7 Modified leaves Reproductive leaves Bracts

Plant Growth Plant Growth: 1) Indeterminate: Grow throughout life 2) Growth at “tips” (length) and at “hips” (girth) Growth patterns in plant: 1) Meristem Cells: Dividing Cells 2) Differentiated Cells: Cells specialized in structure & role Form stable, permanent part of plant

Plant Growth 1) Primary Growth: Apical Meristems: girth length 1) Primary Growth: Apical Meristems: Mitotic cells at “tips” of roots / stems 1) Increased length 2) Specialized structures (e.g. fruits) 2) Secondary Growth: Lateral Meristems: Mitotic cells “hips” of plant Responsible for increases in stem/root diameter

Plant Growth Shoot apical meristem Leaf primordia Young leaf Developing vascular strand Figure 35.16 The shoot tip Axillary bud meristems

Plant Growth Two lateral meristems: vascular cambium and cork cambium Shoot tip (shoot apical meristem and young leaves) Lateral meristems: Axillary bud meristem Vascular cambium Cork cambium Root apical meristems Primary growth in stems Epidermis Cortex Primary phloem Primary xylem Pith Secondary growth in stems Periderm Cork cambium Primary phloem Secondary xylem Figure 35.11 An overview of primary and secondary growth

Plant Growth Stem – Secondary Growth: thicker, stronger stems primary phloem thicker, stronger stems Vascular Cambium: between primary xylem and phloem vascular cambium primary xylem epidermis Produces inside stem: pith A) Secondary xylem moves H2O, inward B) Secondary phloem moves sugars, outward cortex primary xylem dividing vascular cambium primary phloem

Plant Growth Vascular Cambium: Secondary growth primary phloem dividing vascular cambium new secondary xylem secondary phloem primary phloem vascular cambium primary xylem secondary xylem pith cortex Secondary growth Vascular cambium Growth Secondary xylem After one year of growth After two years phloem Vascular cambium X P C

Plant Growth Stem – Secondary Growth: Cork Cambium: ring Vascular ray Secondary xylem Heartwood Sapwood Bark Vascular cambium Secondary phloem Layers of periderm Located under outer surface; produces periderm Dead at maturity Protection

Plant Growth Stem – Secondary Growth: heartwood (xylem) sapwood vascular cambium phloem annual ring Sapwood = Young xylem, water Heartwood = Old xylem, support Seasonal Growth = annual rings Secondary phloem = grows outward older phloem crushed late xylem early xylem

Plant Growth RESULTS 2 1.5 Ring-width indexes 1 0.5 1600 1700 1800 Figure 35.21 1600 1700 1800 1900 2000 Year Using dendrochronology to study climate

Plant Growth Living tree or dead tree? Figure 35.23 Is this tree living or dead?

A cross section of what tissue is pictured? A. Monocot root B. Dicot root C. Monocot stem D. Dicot stem

Things To Do After Lecture 5… Reading and Preparation: Re-read today’s lecture, highlight all vocabulary you do not understand, and look up terms. Ch. 35 Self-Quiz: #1, 3, 6, 7 (correct answers in back of book) Read chapter 35, focus on material covered in lecture (terms, concepts, and figures!) Skim next lecture. “HOMEWORK” (NOT COLLECTED – but things to think about for studying): Compare and contrast monocots and dicots. List the different types of plant cells and describe which tissues and organs they make up, including roles for each organ. Explain the different between apical and lateral meristems and how growth occurs. Discuss the composition of bark and it’s function for plants (do all plants have this tissue?)