Monday’s Topics 1.Overview of photosynthesis 2.Key early photosynthesis experiments 3. Photosynthesis: light reactions.

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Presentation transcript:

Monday’s Topics 1.Overview of photosynthesis 2.Key early photosynthesis experiments 3. Photosynthesis: light reactions

From which molecule does the O2 released by plants come from? What happens to H20 during photosynthesis? (more than one answer might be correct) a. Oxidized b. Reduced c. Activated d. Looses electrons e. Gains electrons Which are the energy rich compounds generated during the light reactions?

Today’s Topics 1.Electron transfer during the light reactions 2.Regulation of photosynthesis 3.Evolution of photosynthetic systems

Energy H20H20 2 H + + ½ O 2 The Electron Transfer Process

Oxidized State Reduced State PSI Light PSI + PSII Light PSI Light Darkness Time Duysens & Amesz (1962) Cytochrome

Z-Scheme: Electron transfers (redox reactions)

Photosynthesis: Light Reactions

Reactions of Plastoquinones

Photosynthesis: Light Reactions

Electron Transfer in the Cytochrome b6f Complex

Photosynthesis: Light Reactions

Herbicides and the study of photosynthesis Blocks electron transfer to plastoquinone Blocks electron transfer in PSI

Excercises What will the status of the plastoquinone pool in DCMU treated plants? a. Reduced b. Oxidiced What will the status of the plastocyanin pool be in Paraquot treated plants? a. Reduced b. Oxidiced

Photosynthesis: Light Reactions

Today’s Topics 1.Electron transfer during the light reactions 2.Regulation of photosynthesis 3.Evolution of photosynthetic systems

Problems with the electron transfer process A.Excess energy absorbance B.Electron balances between PSI and PSII

Energy H20H20 2 H + + ½ O 2 Problems with the electron transfer process

PHOTOPROTECTION

Nonphotochemical Quenching a.Reorganization of antenna complexes b.Special carotenoids: xantophylls c.Thylakoid lumen pH

Problems with the electron transfer process A.Excess energy absorbance B.Electron balances between PSI and PSII

Energy partitioning between photosystems LHCIILHCII-P

Topics 1.Electron transfer during the light reactions 2.Regulation of photosynthesis 3.Evolution of photosynthetic systems

How does a plant make more chloroplasts? Where do the chloroplasts come from? Do all chloroplasts in all algae and plants have the same origin? Where are chloroplast proteins made?

What are the main open questions about the origin of oxygenic photosynthesis? Which are the current hypothesis to explain the beginning of oxygenic photosynthesis?

Next lecture: -Carbon reactions of photosynthesis Light Reactions ATP NADPH Carbon Reactions H 2 0 O 2 CO 2 CH 2 0