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Presentation transcript:

Pg. 367

Membrane Protein Movement Tethered to other Membrane proteins Single Protein Tethered to cytoskeleton Pg. 385

Cytosol Nucleus Mitochondria Chloroplasts ER Golgi Endosome Lysosome Transmembrane Secreted

Cytosol Nucleus Mitochondria Chloroplasts ER Golgi Endosome Lysosome Transmembrane Secreted

The Nucleus

Nuclear Structure

Nuclear Lamina

Nuclear Structure

Nuclear Pore Complex

Protein Transport to the Nucleus Nuclear Localization Signal (NLS) -Pro-Pro-Lys-Lys-Lys-Arg-Lys-Val- N C Table 15-1 (page 504)

NLS is Necessary for Nuclear Import Protein with NLS Nucleus Nuclear protein with NLS removed from protein Pg. 504

NLS is Sufficient for Nuclear Import Nucleus Nuclear protein with NLS removed from protein Cytosolic Protein Without NLS With NLS Pg. 504

Nuclear Import Pathway

Pg. 541

Regulated Nuclear Import NLS “masked” Allostery! NLS “exposed” Pg. 542

Cytosol Nucleus Mitochondria Chloroplasts ER Golgi Endosome Lysosome Transmembrane Secreted

ER Import Sequence Table 15-1 (page 504) Recognition Sequence N-Met-Met-Ser-Phe-Val-Ser-Leu-Leu-Leu-Val-Gly-Ile-Ile-Phe-Trp-Ala-Thr- C Hydrolysis of this peptide bond by Signal Peptidase enzyme Table 15-1 (page 504)

N

Membrane “budding” N C N C N C N C N C

Clathrin “triskelion”

Clathrin “Cage”

Dynamin GTP Hydrolysis

Vesicle Fusion with Membrane

v-SNARE/t-SNARE Interaction

Cytosol Outside of cell Membrane fusion step is regulated Secretory vesicle Ca++ Outside of cell

Botulism Toxin Clostridium botulinum Protease

Botulism Toxin t-SNARE (green) Botulism Toxin (tan)

Botulism Toxin Botulism Toxin Protease