Kingdom Plantae 1.Alternation of generations. 2.Haploid gametophyte that produces gametes by mitosis 3.Diploid sporophytes that produce spores by meiosis 4.mostly autotrophic 5.Fertilization by sperm or pollen nucleus produces a diploid embryo, which is kept within female sex organ on parent plant. 6.First known from the mid-Ordovician, some 476 million years ago.

Land Plants There are four main groups of land plants: 1.Bryophytes, including mosses, liverworts and hornworts. 2.Pteridophytes, including ferns and seedless vascular plants. 3.Gymnosperms or conifers. 4.Angiosperms or flowering plants.

CLASSIFICATION OF SEEDLESS VASCULAR PLANTS There are two phyla of pteridophytes found in the modern flora: Licophyta and Pterophyta. 1. Phylum Lycophyta. There are about 15 genera of lycophytes and approximately 1000 living species. This phylum includes the Lycopods (club mosses), Selaginella (spike moss) and Isoetes (quillwort). This evolutionary line extends back into the Devonian ( mya) but were most prevalent in the wet swamps of the Carboniferous period ( mya).

Lycophyta The Lycophyta eventually split up into two evolutionary lines. 1.The first were very large woody trees that did not survive in the drier climate at the end of and after the Carboniferous age. In the Carboniferous some lycophytes were forest- forming trees more than 35 meters tall. 2.The second and the surviving group of Lycopods are the small and herbaceous trees.

CHARACTERISTICS OF THE LYCOPHYTA Lycophyta are characterized by 1.microphyllous leaves, 2.a special spore producing body called a strobilus, 3.the presence of true vascular stems, roots and leaves.

Importance of the Lycophyta Lycophyta remains became the largest coal deposits of all geologic time.

Some important terms: The sporophytes of lycophytes consist of true roots, stems and leaves. Sporophylls are specialized leaves that bear sporangia and are organized into a structure called the strobilus (pl. strobili). Some Selaginella are heterosporous; Lycopodium is homosporous.

SELAGINELLA There are about 700 species of Selaginella, showing a wide range of characters. There are about 38 species in North America. Selaginellas are mostly tropical and subtropical. The genus is overdue for a revision which might include subdivision into several genera.

SELAGINELLA The gametophyte that arises from the microspore is called a microgametophyte. The microgametophyte never leaves the wall of the microspore; it is endosporic. It is not photosynthetic but heterotrophic; its supply of nutrients is limited to what is contained in the original microspore. Its development is limited to the formation of an antheridium (sterile jacket) containing up to 32 sperm. Thus, the microgametophyte can be thought of as a male structure. The microspore wall and sterile jacket rupture in free water to release the sperm to swim to the egg.

Megagametophyte The gametophyte that arises from the megaspore is called a megagametophyte. It too is endosporic and heterotrophic. Its large volume means that it can contain considerable reserves....usually oils. After cracking the megaspore wall, the megaspore divides into several sterile cells that hold the nutrients, some rhizoid cells that protrude from the cracks, and a few archegonia with eggs.