9.Almost Rosids to Rosids

Santalales Lots of parasitism Polyacetylenes 1-seeded ovules, pendulous and free- central placentation Molecules In ours, flowers not prominent Families not on required list

“Santalaceae” Hemiparasites Root parasites

Viscaceae Epiphytic parasites Haustoria Stems jointed Lvs opp, oft. Coriaceous or succ. Flrs unisex, inconspic, radial T3-4A3 (no fil),G3-4, inf. Viscous berry Passive gametophyte Pantrop + some temperate

Loranthaceae Us. Epiphytes Lv opp or almost so Flrs bisex, rad or bilat Ca rimCo(3)5-6(9), distinct or conn, oft. Red or yellow, A=Co, opp, fil adnate Aggr. gametophyte Tropical

ROSIDS Weakly supported group Also some lack of clarity of relationships within Lots of representatives in our part of the world Hypanthium features in many families Radial symmetry common

Saxifragales Some families chiefly (only?) on molecular data Hypanthium, 2-united carpels, glandular leaf teeth Retained: 5-merous flowers with separate parts

Saxifragaceae Two interpretations Shaw – as in text; Welsh – much broader (older) Herbs, often basal leaves, alt. NO STIPULES Flr rad to bilat, HYPANTH, Ca4-5 sometimes connate, Co4-5, oft. Clawed, St G2(-5), connate, varying position, axile, oft. nectar disk Fr. Capsule or follicle

Crassulaceae

Geraniales Five families, 1 in north Utah Morphological features of order  Leaf margins with glandular teeth  Inflorescence cymose  Calyx persistent in fruit  A few other characters  Anomocytic stomates  Vessel elements with simp[le perforations

Geraniaceae Remarkable feature is fruit Herbs to subshrubs; stems usually jointed at nodes Hairs glandular, with aromatic oils (I.e., smelly plants) Stipules present, leaves with gland. Teeth K 5 C 5 A (5)10,15, filaments often slightly connate G 5conn Fruit a schizocarp, persistent style column

Note the fruits in picture above and on the right Persistent style

Malpighiales This order contains 7.8% of the diversity of angiosperm diversity There does not seem a whole lot tying it together, see: The Angiosperm Phylogeny Group’s home page

Euphorbiaceae