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Entomology By Dr. Fan dong
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Insect Thorax The thorax consists of three segments - the prothorax (front), the mesothorax (middle) and the metathorax (back). Each of these segments bears a pair of jointed legs.
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In winged insects, the meso- and metathoracic segments each bear a pair of wings--the forewings and hindwings forewings hindwings
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In many insects one or both pairs of wings have been lost, or modified into other structures, during the course of evolution.
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Construction of thorax
Prothorax: Pronotum Propleuron: pleural sulcus-episternum and epimeron prosternum
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Construction of Pterothorax
Notum: three sulcus: antecostal sulcus, prescutal sulcus, scutoscutellar sulcus Acrotergite, prescutum, scutum, scutellum Anterior notal wing processes and posterior notal wing processes
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scutellum
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Construction of Pterothorax
Pleuron : pleural sulcus-episternum and epimeron Sternum : two sulcus: presternal sulcus and sternacostal sulcus Eusternum (presternum , basisternum , sternellum) and intersternite
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Thoracic legs coxa trochanter femur tibia tarsus pretarsus: claws
pulvillus arolium
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Mosquito’s femur (2070x)
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Drosophila: claw and pulvillus
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Types of thoracic legs Walking legs : ambulatorial legs
Jumping legs : saltatorial legs Grasping legs : raptorial legs Digging legs : fossorial legs Swimming legs : natatorial legs Clasping legs Pollen-carrying legs : corbiculate legs Clinging legs : scansorial legs
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Walking legs
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Digging legs
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Jumping legs
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Grasping legs
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Pollen-carrying legs
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Clasping legs
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Swimming legs
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Clinging legs
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Insect wings
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Functional wings exist during the adult stage
Membranous, parchment-like, or heavily sclerotized Most insects have two pairs of wings -- one pair on the mesothorax and one pair on the metathorax
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Purpose of Flight in Insects
Search for food or a resting habitat Escape from enemies and search actively for mates Be able to spread over the globe; if conditions became unfavorable in one place, they took to the air to find another
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Function of wings serve as organs of flight
protective covers (Coleoptera and Dermaptera) thermal collectors (Lepidoptera) gyroscopic stabilizers (Diptera) sound producers (Orthoptera) visual cues for species recognition and sexual contact (Lepidoptera).
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In most cases, a characteristic network of veins runs throughout the wing tissue.
These veins are extensions of the body's circulatory system. They are filled with hemolymph and contain a tracheal tube and a nerve. In membranous wings, the veins provide strength and reinforcement during flight. Wing shape, texture, and venation are quite distinctive among the insect taxa and therefore highly useful as aides for identification.
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Structure of insect wing
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Apical angle of a mosquito (80x)
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Special wing modifications include the structures called halteres found in the Diptera (true flies), and the structures called elytra (singular: elytron) found in the Coleoptera (beetles) and Dermaptera (earwigs):
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Membranous wings: such wings are very thin and like cellophane
Types of insect wings Membranous wings: such wings are very thin and like cellophane
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Elytra -- hard, sclerotized front wings that serve as protective covers for membranous hind wings
Coleoptera
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Hemelytra -- front wings that are leathery or parchment-like at the base and membranous near the tip
Hemiptera
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Tegmina -- front wings that are completely leathery or parchment-like in texture
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lepidotic wings: Scaly wings -- front and hind wings covered with flattened setae (scales)
Lepidoptera
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Piliferous wings: Hairy wings -- front and hind wings clothed with setae
Trichoptera
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Fringed wings -- slender front and hind wings with long fringes of hair
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Halteres -- small, club-like hind wings that serve as gyroscopic stabilizers during flight
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Halteres
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front wings and hind wings
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Jugate form
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Frenulum -- Bristle near base of hind wing that holds front and hind wings together
Lepidoptera
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amplexi form
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Hamuli -- tiny hooks on hind wing that hold front and hind wings together
Hymenoptera
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fold form
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Wing Venation Insect contained 7 longitudinal veins.
These veins (and their branches) are named according to a system devised by John Comstock and George Needham -- the Comstock-Needham System
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longitudinal veins Costa (C) -- the leading edge of the wing
Subcosta (Sc) -- second longitudinal vein (behind the costa), typically unbranched Radius (R) -- third longitudinal vein, one to five branches reach the wing margin Media (M) -- fourth longitudinal vein, one to four branches reach the wing margin Cubitus (Cu) -- fifth longitudinal vein, one to three branches reach the wing margin Anal veins (A1, A2, A3) -- unbranched veins behind the cubitus Jugal veins ( J )
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Crossveins crossveins abbreviation longitudinal veins ligated
humeral crossvein radial crossvein sectorail crossvein radiomedial crossvein medial crossvein mediocubital crossvein h r s r-m m m-cu C/Sc R1/R2 R5/R4, or R2+3/R4+5 R4+5/M1+2 M2/M3 M3+4/Cu1
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Variation of veins Increase:accessory veins –R1a, R1b
intercalary veins—IM1 Decrease: combination
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Cells Cell Enclosed cell Open cell Open cell Enclosed cell
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Veins and cells
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Articulation Articulation is an important condition for wing’s all sort of activities. At the end of wings there are some ossicles as joints-pteralia Humeral plate Axillary plates First axillary plates; second axillary plates; third axillary plates Median plates Inner median plate Outer median plate tegula
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Wings’ placement When insect is resting , wings’placement have three types: Open and flat: dragonfly ; some moths; mosquitoes Upright on the back: mayfly ; some butterflies Fore wings cover the back wings: beetles; bugs,grasshoppers
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Flying Wing’s movement: up and down; back and forth
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