1. G.R. Wiggans Animal Improvement Programs Laboratory Agricultural Research Service, USDA Beltsville, MD george.wiggans@ars.usda.gov G.R. WiggansCouncil meeting April 2010 (1) Improved reliability of genomic evaluations

2. G.R. Wiggans Council meeting April 2010 (2) What is reliability l Squared correlation of PTA with true breeding value l Increases with amount of relevant information l Initially derived from the inverse of coefficient matrix l Approximation needed as number of genotyped animals got large

3. G.R. Wiggans Council meeting April 2010 (3) Change implemented in April l Initial approximation assumed all animals received the same amount of information from genomics l Genomic relationship of each animal with the predictor animals routinely calculated l These relationships weighted by reliability and summed now are basis for calculating contribution of genomics to Reliability

4. G.R. Wiggans Council meeting April 2010 (4) Reliability for young HO Bulls 0 500 1000 1500 2000 2500 3000 3500 4000 52535455565758596061626364656667686970717273747576777879 Milk REL Number of Bulls N = 15,226

5. G.R. Wiggans Council meeting April 2010 (5) Reliabilities for HO born ≥ 2005 No Traditional EvaluationWith Traditional Evaluation TraitMaleFemaleMaleFemale N1522675367523191 Milk (lb)73.973.785.877.9 Protein (lb)73.973.785.877.8 PL (mo)64.063.670.167.0 SCS69.769.578.173.0 DPR (%)61.661.266.564.6 PTAT70.470.178.374.5 Sire CE64.961.780.863.5 Daughter CE60.259.069.561.8 Sire SB59.858.766.259.6 Daughter SB58.357.664.959.6 Net Merit ($)68.668.377.872.0

6. G.R. Wiggans Council meeting April 2010 (6) Bulls First Traditional Eval. Jan., 2010 Genomic AugustJanuary TraitNPTARELPTARELDiff PTA Milk70353376.347583.4-58 Protein70317.676.317.083.6-0.6 PTAT4711.070.41.077.50.0 Traditional AugustJanuary TraitNPARELPTARELDiff PTA Milk70365742.047874.9-179 Protein70322.142.017.074.9-5.1 PTAT4711.040.90.965.1-0.1

7. G.R. Wiggans Council meeting April 2010 (7) Cows First Traditional Eval. Jan., 2010 Genomic AugustJanuary TraitNPTARELPTARELDiff PTA Milk59477274.484575.473 Protein57025.574.429.475.13.9 PTAT2941.769.41.871.60.1 Traditional AugustJanuary TraitNPARELPTARELDiff PTA Milk59477435.694351.1167 Protein57026.235.232.949.86.7 PTAT2941.636.81.748.90.1

8. G.R. Wiggans Council meeting April 2010 (8) Adapting to more genotypes l Calculating Genomic Matrix may be too time consuming with many 3K genotypes l May sum the allele content of predictor population l Reliability then based on sum of reliabilities of trad PTA and fraction of alleles in common with predictor population

9. G.R. Wiggans Council meeting April 2010 (9) Conclusions l Contribution of genomics to reliability calculated individually l Based on relationship to predictor population and their Rel of Trad PTA l Mode Rel is 74-75 for bulls without a trad evaluation