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Phylum Mollusca
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Mollusc Characteristics
90,000+ living species 70,000+ fossil species Mantle Pair of dorsal folds of the body wall Forms a mantle cavity Modified into gills or lungs Secretes shell Coelom Limited to area around heart
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Mollusc Characteristics (continued)
Muscular Foot Variously adapted for locomotion Radula Odontophore - supporting cartilage of the radula
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Mollusc Characteristics (continued)
Open Circulatory System Trocophore larva Veliger sometimes follows trocophore
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Class Polyplacophora Chitons ~ 1000 species 7 or 8 dorsal plates
Numerous gills found in the mantle cavity
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Chiton Anatomy
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Class Scaphopoda Tusk Shells ~ 900 species Tubular tusk shaped shell
Captacula - mucus-covered, ciliated tentacles
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Scaphopod Anatomy
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Class Gastropoda Snails, slugs, limpets, whelks, conchs, periwinkles, sea slugs, sea hares, and sea butterflies ~ 70,000 species Well developed head Large flat foot Torsion - during development some body parts rotate relative to the mouth and head Coiling - the shell is coiled and typically conispiral
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Gastropod Anatomy
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Gastropod Torsion
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Evolution of Shell Coiling in Gastropods
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Conispiral shells are either right-handed (dextral) or left-handed (sinistral)
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Figure: 16_34b Title: Gastropod mollusks Caption: (b) Spanish shawl sea slugs prepare to mate.
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Gastropod Eggs
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Class Bivalvia Mussels, Clams, Scallops, Oysters, and Shipworms
~ 15,000 species Two-lobed mantle Two valves Hatchet shaped foot Head is greatly reduced Most are filter feeders
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Bivalve Anatomy
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Bivalve Gills
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Bivalve Hinge Ligament and Adductors
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Bivalve Development
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Glochidium Larvae of Freshwater Clams
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Figure: 16_35b Title: Bivalve mollusks Caption: (b) Mussels attach to rocks in dense aggregations exposed at low tide. White barnacles are attached to the mussel shells and surrounding rock.
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Class Cephalopoda Nautiluses, Ammonoids (extinct), Squids, Cuttlefishes, and Octopuses ~ 700 species Well-developed head with a brain and complex eye Foot modified into funnel and arms/tentacles Shell reduced or absent Ink glands in all except Nautiloids Chromotophores present
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Nautilus Anatomy
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Squid Anatomy
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Figure: 16_36b Title: Cephalopod mollusks Caption: (b) The squid moves by contracting its mantle to generate jet propulsion, which pushes the animal backward through the water.
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Cephalopod Eyes
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Figure: 16_36a Title: Cephalopod mollusks Caption: (a) An octopus can crawl rapidly by using its eight suckered tentacles. It can alter its color and skin texture to blend with its surroundings. In emergencies this mollusk can jet backward by vigorously contracting its mantle.
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