1.  Dynamic Energy Budget Theory Tânia Sousa with contributions from : Gonçalo Marques and Bas Kooijman

2.  DEB Theory on Parameter Values

3.   “A comparison of the energetics of different species, ranging from bacteria to whales is reduced in DEB theory to a comparison of sets of different parameters” DEB Theory on Parameter Values

4.  How to obtain DEB parameters?

5.   Life-stages:  EggLarvae (V1 morph?)Juvenile Adult  Growth curvesSpawning season How to obtain DEB parameters?

6.  Kleiber’s Law  Metabolism (respiration or heat production) as a function of mass  Metabolism increases with weight raised to the power 3/4  Max Kleiber originally formulated this basic relationship back in the 1930s. What is the relationship between specific metabolism and weight?

7.   Relationship between specific metabolism and weight? Kleiber’s Law

8.  DEB Theory on Parameter Values: Scales of Life 1 – Blue whale 2 – T-Rex 13 – Komodo dragon 16 – Cyanea (jelly fish) 24 - Largest flower 26 – sequoia Etruscan shrew Brookesia Micra Chameleon

9.   Constant Primary Parameters DEB Theory on Parameter Values

10.   Empirical support: Cells are very similar independent of size of the organism

11.   Design Primary Parameters: Theory on Parameter Values

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13.   It allows us to make a first rough estimation of DEB parameters knowing L m and the parameters of a reference species Theory on Parameter Values

14.  Kooijman 1986 J. Theor. Biol. 121: 269-282 What are the values for DEB parameters for an animal with L m =1 m? Parameters for a reference animal with L m =1cm and T REF =293 K

15.  Theory on Parameter Values: Flows

16.  Scaling relationships Pink (L sp = 53 cm) Sockeye (L sp = 55 cm) Coho (L sp = 64 cm) Chum (L sp = 68 cm) Chinook (L sp = 87 cm) z = 1 z = 68/87 = 0.78 z = 64/87 = 0.74 z = 53/87 = 0.6 z = 55/87 = 0.63 (Quinn, 2005)

17.  Theory on Parameter Values Kooijman 1986 J. Theor. Biol. 121: 269-282 Something missing? Parameters for a reference animal with L m =1cm What are the values for DEB parameters for an animal with L m =1 m?

18.  Theory on Parameter Values Kooijman 1986 J. Theor. Biol. 121: 269-282 Parameters for a reference animal with L m =1cm and T REF =293 K What are the values for DEB parameters for an animal with L m =1 m and T=308K?

19.  Theory on Parameter Values

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22.   Interspecies comparisons are done for:  Fully grown organism  Abundant food f(X)=1  Null heating length L T =0 Theory on Parameter Values: Flows

23.   Interspecies comparisons are done for:  Fully grown organism  Abundant food f(X)=1  Null heating length L T =0  How do feeding and reproduction rates depend on L m for related species? Theory on Parameter Values: Flows ?

24.   Interspecies comparisons are done for:  Fully grown organism  Abundant food f(X)=1  Null heating length L T =0  How do feeding and reproduction rates depend on L m for related species? Theory on Parameter Values: Flows ?

25.   Interspecies comparisons are done for:  Fully grown organism  Abundant food f(X)=1  Null heating length L T =0  How do feeding and reproduction rates depend on L m for related species? Theory on Parameter Values: Flows

26.   How do feeding and reproduction rates depend on L for the same species? Theory on Parameter Values: Flows

27.   How do feeding and reproduction rates depend on L for the same species? Theory on Parameter Values: Flows

28.  Why does size matter?

29.   Energetics depend on parameter values and parameter values depend on size Why does size matter?

30.   Energetics depend on parameter values and parameter values depend on size  What else matters? Why does size matter?

31.   Von Bertallanffy growth rate DEB Body Size Scaling Relations

32.   Metabolic rate (measured by O 2 or heat production) DEB Body Size Scaling Relations

33.  Exercises

34. 