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Bivalves and Cephalopods
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one thousand species live in fresh water Zebra Mussel
About one thousand species live in fresh water Zebra Mussel Zebras on native clams (Unionidae)
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Best understood in the context of their
Best understood in the context of their largely sedentary adult life style
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umbo ctenidium Foot Adductor muscle Right valve Labial palps mouth
Excurrent Incurrent siphons mouth Foot Mantle edge
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distinguished by their Ctenidia
Two principal types, distinguished by their Ctenidia and mode of feeding Class Bivalvia Protobranchia Llamellibranchia Filibranchia Eullamellibranchia
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Ctenidia of Bivalves are modified for feeding
except in one group: the Protobranchs
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In most bivalves ctenidia are sheet-like
with high S.A., and are used for feeding
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Captured particles are moved to a ciliated ventral groove and then as a mucus strand to the mouth.
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Summary of Water Flow and Feeding
The sheetlike, arrangement of ctenidia in most bivalves is known as the Llamellibranch condition
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There are two basic type of
Llamellibranch ctenidia in bivalves Filibranch (in mussels) Eullamellibranch (most clams)
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Living Cephalopods Squid Nautilus
Cuttlefish Octopods --The class as a whole is adapted for swimming, suspended in the water by buoyancy mechanisms. -- all are carnivores, occupying the same niche as fishes
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Architeuthys Mesonycheteuthis Colossal squid Dosidicus
Giant squid up to 20 m long Mesonycheteuthis Colossal squid up to 14 m major prey item of sperm whales Dosidicus Humboldt squid, smaller but numerous (2 m) and comes to surface to feed
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Ammonites Largest is 3 m in diameter
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Sub Class Nautiloidea 5-6 species remain from diversity in geologic past Live in the last chamber of a multi-chambered shell Move by forcing water through a siphon.
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Sub Class Nautiloidea How does a Nautilus adjust and maintain its buoyancy in water?
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Cuttles, Squids and Octopuses
tendency is reduction of shell. compensated for by selective ionic control; accumulate ammonium other adaptations to shell loss and life style
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cuttle chameleons of the sea most specialized, agile and cryptic octopus
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Specifically, compensation takes the following forms:
Stealth and other defenses Acute sensory structures Speed Cunning
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A. Stealth and other defenses
Ink Sack : ink with melanin and mucus also dopamine, L-DOPA, tyrosinase L-Dopa is a precursor to catecholamines including dopamine which are involved in neurotransmitter function Tyrosinase is an oxidizing enzyme Pseudomorphs
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A. Stealth and other defenses
Chromatophores: each cell contains a different pigment which may be red, orange, brown, black, yellow or blue
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Cephalopod Chromatophore
The pigments in chromatophores can be black, brown, red, orange or yellow. They are not responsible for producing the blue and green colors seen in some species. Those are produced by iridiophores which are reflective cells. Cephalopods have several organs that allow them to hear high frequency sounds Black, brown, orange or yellow colors Also iridiophore reflective cells Most cephalopods are completely colorblind neurophilosophy.files.wordpress.com
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B. Acute sensory structures
-- Vision -- Lateral line analogs
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C. Speed Rival fish in ability to swim, attaining
burst speeds of 5-10 m second sustained for short distances at speeds of up to 40 km hr. Burst speed is achieved by jet propulsion More gradual swimming is fin undulation
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complex nerve control of locomotion
C. Speed : complex nerve control of locomotion Assures synchronized contraction of mantle muscles
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C. Speed : circulatory system
Hearts Requires a closed, elaborate circulatory system
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D. Cunning -- Relative to body size, the brain is larger than that of
most fish
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Experiments with Octopus vulgaris show their ability to learn.
They learn to choose colors not associated with shock. Observers choose properly colored objects after watching demonstrators
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