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Lotic vs. Lentic Systems
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Comparisons Surface area - 1/18th that of lakes
Age - most stream channels ancient compared to most lakes Flow - the most influence on making stream environments different than lakes
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Flow Produces structure of river, bed, influences biota
Currents vary seasonally, longitudinally, vertically
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Flow Generally <3 m/sec (up to 8 m/sec)
Maximal during greatest runoff (wetter season)
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Flow Current speed decreases as depth increases
Decrease from mid-channel to edges (frictional forces between water, substrate - produce boundary layer)
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Flow Average velocity at 0.6 depth for shallow streams (or average of readings at 0.2 and 0.8 depth in deeper streams or rivers)
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Flow Velocity generally decreases from headwaters to mouth
Headwaters - erosional - larger substrates Near mouth - depositional - smaller substrates
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Flow General pattern not always present
Velocity may be constant, or may increase
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Alternating Habitats Riffles and pools Faster currents in riffles
Regular spacing - riffles 5-7 stream widths apart
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Water Temperatures More rapid changes than in lentic waters
Daily cycles up to 6º C in summer Average temperatures tend to increase downstream
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Water Temperatures Little stratification because of flows
Warmer temps alter viscosity Faster flow Carry less silt Reduces boundary layer
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Water Temperatures Winter - entire stream declines to 0º C before ice forms Ice formation at any layer Frazil or anchor ice
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Light Penetration Less in moving vs. standing waters
Increased turbidity Increased shading by riparian vegetation
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Chemistry Dependent on drainage basin
General inverse relation between discharge and concentrations of dissolved materials
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Chemistry Little zonation of dissolved materials or gasses
Turbulent flow, gasses at saturation for temp., pressure Significant changes by activities of living organisms
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Biota Phytoplankton in streams relatively meager Few planktonic forms
Mostly benthic forms
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Biota In larger streams, plankton always present
Planktonic diatoms dominate Selective elimination of other types?
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Biota Most primary production in small streams from periphyton growing on bottom Unusually productive because of nutrient delivery
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Biota Zooplankton numerous only in larger systems
Dominated by rotifers Few crustacean zooplankters
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Biota Benthic invertebrates diverse (especially insects)
Substrate is major controlling factor
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Biota Soft substrate types found in both flowing, standing waters
Hard substrate types often unique Adaptations to cope with flowing water (flattening, hooks, suckers, secretions, ballast)
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Biota Longitudinal rather than depth zonation
Changes in response to temp., current, food availability
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Biota Fish exhibit distinct habitat, longitudinal differences
Those in pools or downstream zones also found in lakes
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Biota Those in headwaters adapted for life in moving water
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