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The History and Development of Evolutionary Theory
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Fossil Fraud In The News
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Charles Darwin Medical Student Later attended Cambridge
Interest in medicine waned Interest in natural history grew Later attended Cambridge Accepted position as naturalist for voyage of discovery on H.M.S. Beagle
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Voyage of the Beagle
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Darwin’s Voyage Initially convinced that species did not change
Observed many phenomenon Fossils of extinct animals Similar geographic areas had different species Began to change his ideas about species change
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Species Group of interbreeding natural populations that are reproductively isolated from other such groups
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Fossil Relatives
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Similar Species, Different Places
Galapagos finch Mainland finch
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Darwin’s Realizations
Organisms of past & present are related Factors play a role in development of diversity Members of a species change slightly when geographically isolated Organisms from islands often resemble those on a nearby mainland
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Change in Earth Over Time
Charles Lyell (1830s) Uniformitarianism Today’s Earth results from gradual change Challenged catastrophism Fossils are similar, but different than living organisms Evidence that change has occurred
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Artificial Selection A clue for Darwin
Breeder can select for desired characters Depends upon natural variation
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Choosing Parents Changes Species
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Population Growth has Limits
Thomas Malthus ( ) Populations can increase exponentially Food supplies increase arithmetically Factors limit population growth
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Darwin’s Associations
Processes of artificial breeding Factors limiting survival Reproduction within natural populations Environmental factors affect natural populations Traits could be naturally selected
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Darwin’s Conclusion Individuals with attributes well suited to their environment are more likely to survive and pass those attributes to offspring
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Adaptations Naturally occurring heritable traits
Favorable to survival and reproduction
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Natural Selection Process by which organisms having adaptive traits reproduce in greater numbers than those without such traits Changes the species with time
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Evolution by Natural Selection
Populations change over time Change is driven by natural selection for best adapted individuals Descent with modification from common ancestors
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Darwin’s Finches Example of natural selection
Found on Galapagos Islands Show change from common ancestor Beaks are adaptations to food source All are thought to be from a single common ancestor from the mainland
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Adaptive Radiation Population changes as it occupies different habitats within a region
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Darwin’s Finches still Evolving
Rosemary & Peter Grant’s work Galapagos island of Daphne Major Documented evolution of beak size Observing evolution in progress
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Evolution Before Darwin
Darwin was not the first to observe evolution Jean-Baptiste Lamark ( ) Inheritance by acquired characteristics Inaccurately concluded change was due to need Correctly observed that species change, incorrectly defined process of change
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Alfred Russel Wallace Independently discovered natural selection
Influenced by same factors as Darwin Naturalist working in Malaysia Jointly presented theory with Darwin
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Darwin’s Book On the Origin of Species by Means of Natural Selection
Published in 1859 Started many debates Evidence continues overwhelming support
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Evidence for Evolution
Fossils Comparative anatomy Comparative embryology Molecular evidence
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Fossils Preserved evidence of past life Soft parts are rare as fossils
Remains Impressions Traces Soft parts are rare as fossils
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Fossil Record Rock layers provide an ordered record
New layers deposited on top of old layers Provides for relative dating
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Fossil Evidence Past living forms can be observed
Change through time can be documented
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Fossil Dating Direct dating methods are available
Example: carbon dating Gives accurate time relationship Shows that time existed for organisms to develop from ancestral forms
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Age of Earth Until Darwin’s time most thought earth was 6,000 to 10,000 years old Earth is know to be about 4.6 billion years old
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Comparative Anatomy Related organisms have same structure as common ancestor Modifications reflect adaptations Homologous structures Same embryological origin Same underlying anatomy, different function
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Vestigial Structures Homologous structure no longer useful
Diminished over time Example: snake’s vestigial leg bone
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Analogous Structures Structures that appear similar
Found in organisms that do not share embryological origin or underlying anatomy
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Convergent Evolution Creates analogous structures
Change in organisms with different ancestors Structures appear similar
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Comparative Embryology
Similarities among embryos point to common ancestors Similar genes with new instructions over time
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Molecular Evidence All life has DNA as hereditary material
Species with similar DNA are closely related
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Phylogeny Evolutionary history of an organism Phylogenetic tree
Depiction of phylogeny Pattern of evolutionary relationships Ernst Haeckel first used tree depiction
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Taxon Named group of organisms
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Phylogenetic Trees Tips represent taxa
Branches represent evolution over time Nodes represent common ancestors
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Methods to Construct Phylogeny
Comparative characters Genetic information
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End chapter 15
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