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“Spine skin” Marine (or estuarine) Water vascular system
Phylum Echinodermata “Spine skin” Marine (or estuarine) Water vascular system Pentaradial symmetry
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Classes: Crinoidea, Stelleroidea, Echinoidea, Holothuroidea
Echinodermata 6500 species living 13,000 from fossils Classes: Crinoidea, Stelleroidea, Echinoidea, Holothuroidea
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Water vascular system Fluid-filled canals that lead to tube feet Sea star: madreporite, stone canal, ring canal, radial canal, ampulla, tube feet
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Water vascular system Tiedemann’s body: Contains phagocytes; remove foreign matter such as bacteria from incoming seawater
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Ambulacral ossicles support ampullae and tube feet
Water vascular system Ambulacral ossicles support ampullae and tube feet Contraction of ampulla moves fluid to tube feet Muscles in tube feet for retracting
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Water vascular system Cilia on inner surface of tube feet circulate water Gas exchange Fluid similar to seawater; contains coelomocytes, proteins, K ions
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Feather stars and sea lilies Oldest of living echinoderms
Class Crinoidea Lily-like Feather stars and sea lilies Oldest of living echinoderms
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Feeding, repro structures at top of stalk
Class Crinoidea Feeding, repro structures at top of stalk Complete digestive system in calyx: mouth – intestine, anus
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Arms have ambulacral groove with mucus-secreting glands adjacent.
Class Crinoidea Arms have ambulacral groove with mucus-secreting glands adjacent. Food particles stick in mucus, flicked to the ambulacral groove, then mouth
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Brittle stars, sea stars
Class Stelleroidea Armed echinoderms Brittle stars, sea stars
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Brittle stars and basket stars
~ 2100 described Joints allow flexibility Tube feet present Sensitive to light – oral surface
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Brittle stars and basket stars
Tube feet through small holes Digestive system mostly in disc – no anus Bursal slits for water exchange
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Brittle stars and basket stars
Deposit feeders, suspension feeders, carnivores, scavengers Many are nocturnal Many live in associations
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Arms not as distinct from disc as in brittle stars
Sea stars ~ 1600 species Arms not as distinct from disc as in brittle stars Move slowly with tube feet Tube feet move individually
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Figure 20_06
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Digestion: lower cardiac stomach digests food
Sea stars Digestion: lower cardiac stomach digests food Upper pyloric stomach for secreting enzymes and absorption
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Pedicellariae: stalked or sessile
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Class Echinoidea: spine-like
Sea urchins, sand dollars < 1000 species
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Class Echinoidea: spine-like
Ossicles form test Complex system of ossicles and muscles for grazing = Aristotle’s lantern
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Spines attach to skeleton – ball and socket joint
Toxins Ossicles flat and joined = inflexible
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Class Echinoidea: spine-like
Tube feet in 5 double rows of plate Pedicellariae – globular forms have toxin
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Feeding and digestion:
Aristotle’s lantern Teeth protruded to scrape algae or consume food Species w/o lantern usually detritivores
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Mouth – esophagus – intestine – anus
WVS
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Class Holothuroidea: sea cucumbers
~ 1200 species
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Ossicles microscopic Multiple shapes
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Tube feet modified into tentacles around mouth
Mostly deposit-feeders, few filter-feeders
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Digestion system: elongated
Mouth – esophagus – stomach – intestine – cloaca – anus WVS – madreporite in coelomic cavity = no outside connection
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Respiratory tree: connects to cloaca – water supply
Expulsion of internal organs
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Echinoderm repro + development:
Some are asexual Most are dioecious Multiple gonads, gametes into seawater = external fert
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Distinctive ciliated larval form in each class
Free-swimming, planktonic Metamorphosis into adult
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Echinoderm NS No brain 3 nerve networks Ectoneural = ring around esophagus: receives sensory input
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