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Insect-pollinated flowers
Often yellow or blue Have a scent Bird-pollinated flowers Often yellow, orange, or red Do not have a strong scent
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Ultraviolet markings on insect-pollinated flowers
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Bat-pollinated flowers
Often have dusky white petals Are scented Wind-pollinated flowers Often have smaller petals or none at all Have neither scent nor nectar Make large amounts of pollen
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Animal pollinators: Archilochus colubris obtains nectar
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Animal pollinators: Leptonycteris curasoae obtains nectar
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Wind pollination
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Coevolution Reciprocal adaptation Caused by two species
Forming interdependent relationship Affecting the course of each other’s evolution E.g., certain showy flowers + bees
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Pollination Fertilization
Transfer of pollen grains from anther to stigma Fertilization Fusion of gametes Occurs after pollination
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Double fertilization In the ovule, egg fuses with first sperm cell
Zygote is formed Zygote develops into a multicellular embryo in the seed
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Double fertilization, cont.
Two polar nuclei fuse with second sperm cell Triploid nutritive tissue (endosperm) is formed
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Mature flowering plant embryo consists of
A radicle A hypocotyl A plumule Cotyledons (one in monocots, two in dicots)
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For use during germination, a mature seed contains both
A young embryo Nutritive tissue (stored in endosperm or cotyledons)
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Ovules Ovaries Structures with the potential to develop into seeds
Structures with the potential to develop into fruits
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Simple fruits Develop from a single pistil consisting of
Either a single carpel Several fused carpels Some are fleshy at maturity Others are dry
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Aggregate fruits Multiple fruits
Develop from a single flower with many separate ovaries Multiple fruits Develop from the ovaries of many flowers growing close together on a common axis
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Dispersal methods of seeds and fruits include
Accessory fruits The major part of the fruit consists of tissue other than ovary tissue Dispersal methods of seeds and fruits include Animals Wind Water Explosive dehiscence
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An accessory fruit
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Plant growth and development are controlled by
Internal factors like location of cell in plant body Causes some genes in cell to be turned off and others to be turned on, thus affecting gene expression during development Environmental factors like changing day length
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Germination and seedling growth
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Germination is the process of seed sprouting
Internal factors affecting germination Maturity of the embryo Presence or absence of chemical inhibitors Presence or absence of hard, thick seed coats
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Germination, cont. External environmental factors affecting germination include requirements for Oxygen Water Temperature Light
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Photoperiodism is any response of plants to the duration and timing of light and dark
In many plants, flowering is a photoperiodic response Short-day plants Long-day plants Intermediate-day plants
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Photo-periodic responses of short-day and long-day plants
Light treatment Photo-periodic responses of short-day and long-day plants Short-day plant Long-day plant
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Photoperiodism, cont. Day neutral plants
Flowering is not affected by photoperiod
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Phytochrome is The photoreceptor in photoperiodism
A family of ~5 blue-green pigments Each type has two forms named by the wavelength of light they absorb Pr Pfr
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Phytochrome, cont. Pfr is the active form, triggering or inhibiting physiological responses such as Flowering Shade avoidance A light requirement for germination
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Phytochrome This pigment occurs in two forms, Pr and Pfr, and readily converts from one to the other. Red light (660 nm) converts Pr to Pfr, and far-red light (730 nm) converts Pfr to Pr
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Circadian rhythm A regular period in the growth or activities of a plant or organism That approximates the 24-hour day Is reset by the rising and setting of the sun
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Circadian rhythm, cont. Examples are
The opening and closing of stomata Sleep movements
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Plant movements occurring in response to external stimuli
Tropisms Directional growth responses (i.e., the direction of growth is dependent on the direction of the stimulus)
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Phototropism Gravitropism Thigmotropism Heliotropism
Response to the direction of light Gravitropism Response to gravity Thigmotropism Response to contact with a solid object Heliotropism Ability of leaves or flowers to track the sun across the sky
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Gravitropism
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Heliotropism
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Plants produce and respond to hormones that
Act as highly specific chemical signals Elicit a variety of responses that regulate Growth Development
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Hormones Are effective in very small concentrations
Functions of some overlap Many physiological activities are regulated by interactions of several at once
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General mechanism of action of plant hormones
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Plant hormones bind to specific receptor proteins in or on target cells
Binding may trigger production of a second messenger (e.g., Ca2+) Second messenger may bind to and activate or inactivate certain enzymes
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Hormones affecting plant growth and development
Auxin is involved in Cell elongation Tropisms Apical dominance Inhibition of axillary buds by the apical meristem
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Isolating auxin from coleoptiles
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Phototropism and the unequal distribution of auxin
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Hormones, cont. Gibberellins are involved in Stem elongation Flowering
Germination
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Gibberellin and stem elongation
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Hormones, cont. Cytokinins Promote cell division and differentiation
Delay senescence Interact with auxin and ethylene in apical dominance Induce cell division in tissue culture
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Auxin-cytokinin interactions
in tissue culture
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Hormones, cont. Ethylene plays a role in Ripening fruits
Apical dominance Leaf abscission Wound response Thigmomorphogenesis Senescence
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Hormones, cont. Abscisic acid is an environmental stress hormone involved in stomatal closure caused both By water stress In seed dormancy
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Abscisic acid and seed germination
Arrows show where some of the kernels have germinated while still on the ear, producing roots
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Salicylic acid Helps defend plants against
Pathogens Insect pests May bind to a cell receptor to switch on genes to Fight infection Promote wound healing
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