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PowerLecture: Chapter 25 Part I
Animal Evolution – The Invertebrates
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Characteristics of Animals
Multicelled Aerobic heterotrophic eukaryotes Most reproduce sexually
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Animal Origins Originated during Precambrian (1.2 billion million years ago)
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Radial Symmetry radial symmetry Fig. 25-5a, p.406
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Bilateral Symmetry dorsal posterior ventral anterior
Fig. 25-5b, p.406
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Animal Tissues Ectoderm Endoderm Mesoderm
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The Gut - 2 types Saclike gut Complete digestive system
One opening for taking in food and expelling waste Complete digestive system 2 Openings; mouth and anus
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Body Cavities - Acoelomate
epidermis gut cavity no body cavity; region between gut and body wall packed with organs Fig. 25-6, p.406
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Body Cavities - Pseudocoel
epidermis gut cavity unlined body cavity (pseudocoel) around gut Fig. 25-6, p.406
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lined body cavity (coelom)
Body Cavities - Coelom gut cavity lined body cavity (coelom) peritoneum Fig. 25-6, p.406
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Segmentation Repeating series of body units
Units may or may not be similar Insects - segments may be fused
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Segmentation Fig. 25-8c, p.407
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Segmentation Fig. 25-8a, p.407
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Fig. 25-7, p.407 sponges cnidarians flatworms annelids mollusks
roundworms arthropods echinoderms chordates coelom reduced coelom reduced coelom lost pseudocoel molting PROTOSTOMES mouth from blastopore radial ancestry, two germ layers DEUTEROSOMES anus from blastopore bilateral, coelomate ancestry, three germ layers true tissues multicelled body Fig. 25-7, p.407
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Deuterostomes Protostomes Fig. 25-2a, p.404
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p.408
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Choanoflagellates Protozoans most closely related to animals
Resemble collar cells (stay tuned for more on this)
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flagellated collar cell
Animal Origins flagellated collar cell Fig. 25-4a, p.405
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p.408
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Sponges - Phylum Porifera
No symmetry, tissues or organs Reproduce sexually Microscopic swimming larvae
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Sponge Structure water out glasslike structural elements amoeboid cell
pore central cavity semifluid matrix flattened surface cells water in flagellum microvilli nucleus Fig , p.409
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Fig. 25-9a, p.408
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Fig. 25-9b, p.408
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Fig. 25-9c, p.408
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p.408
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Phylum Placozoa One living species, Tricoplax adherens
2-layer body, 3 mm across Fig , p.409
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p.408
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Cnidarian Diversity Scyphozoans Anthozoans Hydrozoans Jellyfish
Sea anemones Corals Hydrozoans
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Phylum Cnidaria Nerve net Hydrostatic skeleton Saclike gut nematocysts
capsule’s lid at free surface of epidermal cell trigger barbed thread inside capsule nematocyst Fig , p.410
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mesoglea-filled bell tentacles Fig b, p.410
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outer epithelium (epidermis) inner epithelium (gastrodermis)
Two Main Body Plans Polyp outer epithelium (epidermis) mesoglea (matrix) Medusa inner epithelium (gastrodermis) Fig , p.410
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Fig a1, p.411
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Fig a2, p.411
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Fig b, p.411
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Fig a, p.431
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Obelia Life Cycle (Hydrozoan)
female medusa male medusa reproductive polyp sperm ovum feeding polyp zygote polyp forming planula Fig a, p.411
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Deuterostomes Protostomes
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Flatworms: Phylum Platyhelminthes
Acoelomate, bilateral, cephalized Most hermaphroditic
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Three Classes Turbellarians (Turbellaria) Flukes (Trematoda)
Tapeworms (Cestoda)
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Planarian Organ Systems Digestive
branching gut pharynx; protrudes onto food, then retracts into the body between feedings
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Planarian Organ Systems Neuro-sensory
rudimentary brain (pair of large ganglia in head) pair of nerve cords that have lateral branchings
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Planarian Organ Systems Reproductive
ovary testis oviduct genital pore
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Planarian Organ Systems Excretory
pair of highly branched tubules that adjust water and solute levels in body Fig d, p.412
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flame cell opening of tubule at body surface Fig e, p.412
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flame cell nucleus cilia fluid filters through membrane folds
Fig f, p.412
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Fig , p.413
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Fig. 25-18, p.413 proglottids scolex
a Larvae, each with inverted scolex of future tapeworm, become encysted in intermediate host tissues (e.g., skeletal muscle) b A human, a definitive host, eats infected, undercooked beef which is mainly skeletal muscle d Inside each fertilized egg, an embryonic, larval form develops. Cattle may ingest embryonated eggs or ripe proglottids, and so become intermediate hosts c Each sexually mature proglottid has female and male organs. Ripe proglottids containing fertilized eggs leave host in feces, which may contaminate water and vegetation. Fig , p.413
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proglottids scolex Fig e, p.413
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Deuterostomes Protostomes
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Rotifers Bilateral Cephalized Crown of cilia at head Complete gut
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Fig a, p.416
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one of two ciliated lobes at head end
mouth modified pharynx; internal jawlike elements, salivary glands cluster of nerve cells gastric gland (esophagus behind it) proto-nephridium stomach intestine cloaca (this type is a chamber for digestive and excretory wastes) anus gland that secretes cementing substance for “toe” one of two “toes” Fig b, p.416
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Two Coelomate Lineages
Protostomes Mollusks Annelids Arthropods Deuterostomes Echinoderms Chordates
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Cleavage Patterns Protostome embryo (spiral cleavage)
Deuterostome embryo (radial cleavage)
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Deuterostomes Protostomes
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Annelids: Phylum Annelida
Segmented, coelomate worms Class Polychaeta Class Oligochaeta Class Hirudinea
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Polychaetes Most marine Bristles extend from parapods on each segment
“jaws” toothlike structures Most marine Bristles extend from parapods on each segment Head end is specialized pharynx (everted) antenna palp (food handling) tentacle eyes chemical-sensing pit parapod
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Polychaetes Fig a, p.414
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Fig b, p.414
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Fig c, p.414
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Leeches - Class Hirudinea
Predators and parasites Less obvious segmentation
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before feeding Fig a, p.414
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after feeding Fig b, p.414
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Earthworm - An Oligochaete
No parapodia, few bristles per segment Nerve cord Dorsal blood vessel Circular muscle Coelom Longitudinal muscle Nephridium Seta (retracted) Nerve cord Fig , p.415
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Earthworm Nephridium bladderlike storage region of nephridium
nephridium’s thin loop reabsorbs some solutes, relinquishes them to blood blood vessels body wall funnel (coelomic fluid with waste enters here) external pore (fluid containing wastes discharged here) Fig , p.415
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Earthworm Circulatory System
Hearts Fig , p.415
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Earthworm Digestive System
Coelomic chambers Crop Gizzard Esophagus Pharynx Mouth Fig , p.415
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Earthworm Nervous System
Brain Nerve cord Fig , p.415
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head end Fig g, p.415
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of outwardly similar body segments are bristles used in locomotion
on both sides of outwardly similar body segments are bristles used in locomotion Fig h, p.415
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Old Genes, New Drugs Fig. 25-1, p.402
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Old Genes, New Drugs p.403
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