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Photosynthesis Chapter 10
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Respiration Photosynthesis
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I) Photosynthesis A) Equation:
CO2+H2O C6H12O6 + O2 B) Occurs in the chloroplasts using the enzymes chlorophyll
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C) Broken into 2 parts: 1) Calvin Cycle (light independent reaction): a) occurs in the stroma of chloroplast
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2) Light reaction a) occurs in the thylakoid membrane a1) fyi: stacks of thylakoids are called grana
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D) Fate of the reactants 1) CO2+H2O C6H12O6 + O2
a) using heavy O, they traced where the elements ended up: CO2+H2O C6H12O6 + O2
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II) Properties of light
A) Electromagnetic spectrum a) red is weakest(longest wavelength 750nm) b) violet is strongest(shortest wavelength 380nm)
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B) Absorption spectrum: areas of visible spectrum that is absorbed by a pigment 1) specific wavelengths(photons) are absorbed by a particular pigment
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a) absorbed means it has enough energy to cause an electron to jump from a lower energy level to a higher one.
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a1) with no-where to go the excited electron will fall back to original orbital, emitting light(fluorescence)
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a2) chloroplasts use a primary electron acceptor molecule to trap the excited electron and use it to make NADPH and ATP instead of losing energy as light and heat
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b) Pigments: b1) chlorophyll a: only pigment that passes electrons directly to the primary electron acceptor
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b2) chlorophyll b: pass excited electrons to chlorophyll a
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b3) Carotenoids: yellows and oranges
b3) Carotenoids: yellows and oranges * do absorb light and pass electrons to chlorophyll * serve in photoprotection: absorb excessive light that may damage chlorophyll
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* xanthophylls: contain oxygen * carotenes: pure hydrocarbons
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C) Action spectrum: region of visible spectrum that does the work(measured by O2 release).
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III) The Light reaction
A) Photosystems 1) groups of pigments that pass electrons along to a chlorophyll a molecule, which is directly associated with a primary electron acceptor
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a) Photosystem I(P700) a1) named after wavelength absorbed by the chlorophyll a associated with the electron acceptor
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a2) photosystem II(P680): this cholorphyll a absorbs a wavelength of 680nm
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The Light Reaction: The “Z” scheme with non-cyclic photophosphorylation
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The Light Reaction: Cyclic-photophosphorylation
Purpose:
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B) Chemiosmosis ATP production from the light reaction:
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IV) The Calvin Cycle (light independent reaction)
A) CO2’s go in, 3-carbon sugars(G3P) come out, which are then assembled into glucose and other organic compounds
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1) Driven by the re-use of a 5 carbon sugar called Ribulose bisphosphate
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a) Enzyme called rubisco attaches a CO2 to ribulose bisphosphate, making a 6-C sugar that splits in two, which chugs through the rest of the cycle
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V) Other forms of carbon fixation
A) “Standard”: C3 plants 1) use rubisco in the Calvin cycle, making 3- carbon sugars after the initial carbon fixation phase
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2) dependent on open stomata to provide CO2. No CO2 – no Calvin cycle
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B) C4 Plants 1) Contain a layer of bundle sheath cells between the vascular tissue and the mesophyll
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2) mesophyll cells use the enzyme PEP carboxylase to add CO2 to PEP(3C) to make oxaloacetate(4C).
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3) This changes to Malate(4C) which diffuses into a bundle sheath cell and drops off a CO2, which will then go into the calvin cycle
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a) this keeps CO2 concentration high in the bundle sheath cell so O2 won’t be able to bind rubisco
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C) CAM plants 1) adaptation in succulents and cacti, among other plants
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2) Make organic acids during the night, close their stomata during the day, and cut CO2 from the organic acid to pump into the calvin cycle
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3) compared to C4, C4 plants separate steps spatially, while CAM plants separate the steps temporally.
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