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Population Genetics
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The Gene Pool Members of a species can interbreed & produce fertile offspring Species have a shared gene pool all of the alleles of all individuals in a population
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The Gene Pool Different species do NOT exchange genes by interbreeding
Different species that interbreed often produce sterile or less viable offspring e.g. Mule
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Populations A group of the same species living in an area
No two individuals are exactly alike (variations) More Fit individuals survive & pass on their traits
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The Hardy-Weinberg Principle
Used to describe a non-evolving population. Natural populations are NOT expected to actually be in Hardy-Weinberg equilibrium. Deviation from Hardy-Weinberg equilibrium usually results in evolution Understanding a non-evolving population, helps us to understand how evolution occurs
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5 Assumptions of the H-W Principle
Large population size - small populations have fluctuations in allele frequencies (e.g., fire, storm). No migration - immigrants can change the frequency of an allele by bringing in new alleles to a population. No net mutations - if alleles change from one to another, this will change the frequency of those alleles
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5 Assumptions of the H-W Principle
Random mating - if certain traits are more desirable, then individuals with those traits will be selected and this will not allow for random mixing of alleles. No natural selection - if some individuals survive and reproduce at a higher rate than others, then their offspring will carry those genes and the frequency will change for the next generation.
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Speciation Formation of new species
One species may split into 2 or more species A species may evolve into a new species Requires very long periods of time
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Methods of speciation Allopatric speciation: a physical barrier divides one population into two or more populations Sympatric speciation: a species evolves into two ore more different species within the same environment
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Patterns of evolution that lead to speciation
Adaptive radiation Aka divergent evolution When one species gives rise to many species in response to the creation of new habitat or ecological opportunities (new food source) Often follows mass extinctions Coevolution When the evolution of one species affects the evolution of another species
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Patterns of evolution that lead to speciation
Convergent evolution When unrelated species evolve similar traits even though they live in different parts of the world Similar ecology and climates in different geographical areas Could lead to analogous structures
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5 Causes of evolution 1. Genetic Drift
the change in the gene pool of a small population due to chance Two types: Bottle neck effect Founders effect
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Factors that Cause Genetic Drift
Bottleneck Effect a drastic reduction in population (volcanoes, earthquakes, landslides …) Reduced genetic variation Smaller population may not be able to adapt to new selection pressures
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Loss of Genetic Variation
Cheetahs have little genetic variation in their gene pool This can probably be attributed to a population bottleneck they experienced around 10,000 years ago, barely avoiding extinction at the end of the last ice age
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Founder Effect occurs when a new colony is started by a few members of the original population Reduced genetic variation May lead to speciation
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2. Natural Selection - Cause ADAPTATION of Populations
- success in reproduction based on heritable traits results in selected alleles being passed to relatively more offspring (Darwinian inheritance) - Cause ADAPTATION of Populations
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3 Modes of Natural Selection
Directional Selection Favors individuals at one end of the phenotypic range Most common during times of environmental change or when moving to new habitats Disruptive selection Favors extremes over intermediate phenotypes Occurs when environmental change favors an extreme phenotype
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3 Modes of Natural Selection
Stabilizing Selection Favors intermediate over extreme phenotypes Reduces variation and maintains the curent average Example: Human birth weight
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3. Gene Flow 4. Mutation 5. Non-random mating
-is genetic exchange due to the migration (immigration or emigration) of fertile individuals or gametes between populations 4. Mutation a change in an organism’s DNA Mutations can be transmitted in gametes to offspring 5. Non-random mating Mates are chosen on the basis of the best traits
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Preventing speciation
Prezygotic isolation: mechanisms that prevent reproduction by making fertilization unlikely Prevent new genotypes from entering a gene pool through geographic, ecological and behavioral differences Temporal isolation: timing differences prevent interbreeding Ex: closely related species of fireflies mate at different times of night Behavioral isolation: mating behaviors prevent interbreeding Ex: differences in mating calls between closely related species of birds
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Preventing speciation
Postzygotic isolation: mechanisms that prevent a hybrid (offspring of interbreeding) from surviving or reproducing When two different species reproduce, their offspring are typically infertile Ex: ligers are the product of a male lion and female tiger mating and are sterile Lions are tigers are different species but are close enough to interbreed
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How fast does evolution occur?
Two theories: Gradualism: evolution proceeds in small, gradual steps Punctuated equilibrium: rapid spurts of genetic change cause species to diverge quickly - small periods of change, large periods of no change
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Allopatric speciation Sympatric speciation Adaptive evolution
On pg 110 draw illustrations for the following terms/phrases to help you remember them: Hardy- Weinberg Speciation Allopatric speciation Sympatric speciation Adaptive evolution Coevolution Convergent evolution Genetic drift Founder’s effect Bottleneck effect Directional selection Disruptive selection Stabilizing selection Gene flow Prezygotic isolation Postzygotic isolation
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