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A2E Today: 1. Introduction to Photosynthesis 2. Lunch 3. Guest Speaker (noon; Green Room)
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? PHOTOSYNTHESIS! Brainstorm: 4 Things We Know About Photosynthesis
4 Questions About Photosynthesis Why is Photosynthesis important?? ?
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A Quick Introduction to the Carbon Cycle
What does it mean to be a carbon source? A carbon sink? Examples?
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6CO2 + 6H2O + Light Energy C6H12O6 + 6O2
PHOTOSYNTHESIS Photosynthesis is an enzyme-mediated, metabolic process performed by photoautotrophs 6CO2 + 6H2O + Light Energy C6H12O6 + 6O2 or CO2 + H2O CH2O + O2
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Overview of Leaf and Chloroplast
Anatomy Note the interplay of Form and Function!
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Overview of Photosynthesis
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The LIGHT Reactions
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LIGHT used in Photosynthesis
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LIGHT used in Photosynthesis
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Engelmann figured out the ACTION SPECTRUM in 1883!!
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When PHOTONS hit a Chlorophyll Molecule they EXCITE ELECTRONS
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When PHOTONS hit a Chlorophyll Molecule they EXCITE ELECTRONS
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EXCITED ELECTRONS are passed from molecule to molecule until they reach the REACTION CENTER
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Noncyclic electron flow generates
ATP and NADPH
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What is NADPH? I carry electrons!
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of NONCYCLIC ELECTRON FLOW
Cartoon Schematic of NONCYCLIC ELECTRON FLOW in the LIGHT REACTIONS
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ELECTRON FLOW may also by CYCLIC
Cyclic Electron Flow allows cells to “catch up” on the production of ATP!
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CHEMIOSMOTIC synthesis of ATP
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CHEMIOSMOTIC synthesis of ATP
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Overview of Photosynthesis
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The Light-Independent Reactions
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RUBISCO
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THE DOWNSIDE: Plants must open their
STOMATA to acquire CO2 and release O2. Open Stomata allow for Transpiration (the evaporative loss of water)
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O2 H2O CO2
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How could plants minimize this evaporative loss of water (transpiration)??
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The Pitfall…
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Why is this enzyme so sloppy??
RUBISCO Why is this enzyme so sloppy??
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Photorespiration generates no ATP, NADPH, or glucose!
Photorespiration is a major limiting factor in the production levels of C3 PLANTS Photorespiration generates no ATP, NADPH, or glucose! Rice Up to 50% of the yield of many important agricultural crops like rice, soybeans, and wheat may be lost to photorespiration. Soybeans
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Alternate Strategy #1: the C4 PATHWAY
Carbon fixation and the Calvin Cycle are separated. Fixation occurs in the MESOPHYLL CELLS while the Calvin Cycle occurs in the BUNDLE-SHEATH CELLS. corn This separation allows the plants to minimize photorespiration and maximize sugar production. sugarcane
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An Overview of the C4 Pathway:
Results in a SPATIAL SEPARATION of CO2 acquisition and fixation!
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Alternate Strategy #2: CAM
CAM plants open their STOMATA at night to acquire CO2. This CO2 is used during the day for fixation in the Calvin Cycle. (Temporal separation!)
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C4 and CAM Strategies:
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What Can You Do If You Understand Photosynthesis??
Feed a growing human population?? Maybe! Maybe not!
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