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Phyla Cnidaria and Ctenophora
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Domain Eukarya Kingdom Animalia
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Phyla Cnidaria Marine members include jellyfishes, sea anemones, corals, and hydroids.
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Cnidaria and Ctenophora exhibit radial symmetry
Body Plan Cnidaria and Ctenophora exhibit radial symmetry
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Relatively simple, diffuse nerve net
Nervous System Relatively simple, diffuse nerve net lacks a central brain to process sensory info. or to organize complex responses
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Different degrees of centralization in metazoan brains.
Different degrees of centralization in metazoan brains. (a) Centralized nervous system of an oligochaete worm. (b) Nerve net of a cnidarian polyp representing a typical non-centralized nervous system. Schematized drawings modified with permission from Bullock & Horridge (1965). Arendt D et al. Phil. Trans. R. Soc. B 2008;363: ©2008 by The Royal Society
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Mouth is located at the center of the body on the oral side.
The opposite side is the aboral side
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Box Jellyfish Box Jellyfish
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Uncharacteristically advanced Cnidarian
Uncharacteristically advanced Cnidarian. Primitive eyes (eyespots) are present. When they detect a dark spot, they go the other way.
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Structure and Function
The body may be either vase- shaped (polyp) or bell-shaped (medusa)
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Polyp is sessile and benthic.
Tentacles and mouth face upward
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Medusa is free-swimming
Tentacles and mouth face downward
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All cnidarians have bodies constructed of two layers
Epidermis is outer layer Gastrodermis is inner layer
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A. gastrovascular cavity
B. gastrodermis C. mesoglea D. nematocysts E. epidermis (integument)
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Diploblasty is a condition of the blastula in which there are two primary germ layers: the ectoderm and endoderm.
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Between the layers is a jellylike mesoglea
A centrally located mouth leads to a baglike digestive tract- the gastrovascular cavity
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Cnidarian feeding and Defense
Tentacles that surround the mouth are covered with specialized cells- cnidocytes
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Cnidocytes produce organelles called nematocysts
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Nematocysts are stinging organelles that exit the cnidocyte (cnidoblast) with great force once triggered
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The nematocyst capsule stores a large concentration of calcium ions, which are released from the capsule into the cytoplasm of the cnidocyte when the trigger is activated. This causes a large concentration gradient of calcium across the cnidocyte plasma membrane. The resulting osmotic pressure causes a rapid influx of water into the cell. This increase in water volume in the cytoplasm forces the coiled nematocyst to eject rapidly.
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Some nematocysts are adhesive, others have filaments to entangle prey, and others have spines that puncture prey.
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Tentacles capture prey
Cnidocytes release nematocysts into prey, paralizing it. Tentacles push prey into mouth
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4. Enzymes in gastrovascular cavity break up prey 5
4.Enzymes in gastrovascular cavity break up prey 5.Gastrodermis cells absorb nutrients 6.Waste exits mouth
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Classification of Cnidaria
Four classes Hydrozoa Scyphozoa Cubozoa Anthozoa
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Class Hydrozoa 3700 species Most live as colonial organisms
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Obelia Many polyps attached to branched stalks
Some polyps function in gathering food; others are responsible for reproduction
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Oblia
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Order siphonophores Physalia physalis
Portuguese man-of-war Exists as a colony of medusea and polyps
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Pneumatophore- a single individual polyp (gas- filled float)
Dactylozooids- (tentacles) polyps specialized for capturing prey
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Gastrozooids- polyps specialized for digestion
Gonozooids- polyps specialized for reproduction
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Class Scyphozoa “Cup Animals” Medusa is dominant form of life cycle
Commonly known as jellyfish
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largest (Cyanea capillata, or lion’s mane), which has a bell that measures 6 feet across
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Pulsating motion of the cup propels the jellyfish
Common jellyfish have life cycle that includes both medusa and polyp
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Common jellyfish reproduction
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1. Adult Male and Female
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2. Egg and Sperm are Released
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Fertilization
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Blastula Formation
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Planula Development
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Benthic Attachment
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Polyp Develops
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Stacks of Medusae form and detach
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Young Medusa is Released
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Class Cubozoa Box Jellies Most advanced and deadly eyespot
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(Irukandji, Carukia barnesi) which is the size of your fingernail.
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Class Anthozoa “Flower Animals” ~6100 Species
Examples include Sea anemones and corals
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Sea anemones are benthic polyps
Commonly found in coastal areas Feed on fishes and other small animals
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Symbiotic relationship with clownfish.
Corals are small polyps that usually live in colonies.
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Each polyp cements its calcium carbonate skeleton to the skeletons of others
Live symbiotically with algae
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Phylum Ctenophora “Comb holder”- comb jellies ~100 species
Move by beating cilia (ctenes) lined along body
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Secrete sticky substance from colloblasts.
Some have cnidocytes they recycle from eating cnidarians
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Two tentacles Bioluminescent
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