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Community Ecology Community Community Ecology
all the organisms that live together in a given place Community Ecology study of interactions among all populations in a common environment To answer: In what way do the populations interact?
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Why is the beetle rolling the fecal ball?
Niche An organism’s niche is its ecological role habitat = address vs. niche = job Ex: what it eats, what it lives in, time of day when active, behavior, etc… “How it fits into that ecosystem.” Why is the beetle rolling the fecal ball? Is that frog noctural?
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Competitive Exclusion Principle
No two similar species can occupy the same niche at the same time. Or else….. High tide Competitive Exclusion If Species 2 is removed, then Species 1 will occupy whole tidal zone. But at lower depths Species 2 out-competes Species 1, excluding it from its potential (fundamental) niche. Species 1 Low tide Chthamalus sp. Species 2 Fundamental niches Realized niches Semibalanus sp.
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Symbiotic Relationships
competition (-/-) predation / parasitism (-/+) herbivory (-/+) commensalism (+/0) mutualism (+/+) lichens (algae & fungus) “Freddy fungus and Annie algae took a Lichen to each other!”
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Predation drives evolution
Predators adaptations locate & subdue prey Prey adaptations elude & defend Predation provides a strong selection pressure on both prey & predator horns, speed, coloration spines, thorns, toxins Predation provides a strong selective pressure on prey populations. Any feature that would decrease the probability of capture should be strongly favored.
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Defensive Adapations Mechanical Defense Chemical Defense
Aposematic coloration Cryptic coloration Batesian Mimicry Mullarian Mimicry **All heavily rely on operant conditioning, trial and error learning, in order to be successful. They rely on predators learning what not to eat, and the predator remembering what not to eat.
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Batesian Mimicry Convergent evolution green parrot snake
palatable or harmless species mimics a harmful model green parrot snake Hawkmoth larva puffs up to look like poisonous snake hawkmoth larvae
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Mullerian mimicry cuckoo bee yellow jacket
two or more poisonous species look like each other cuckoo bee yellow jacket - group defense? - predators may evolve innate avoidance
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Coevolution in Community
Predator-prey relationships Parasite-host relationships Flowers & pollinators Long term evolutionary adjustments between species
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Predator/Prey Coevolution…
Biowarfare arms race with garter snakes and newts.
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Community Structure Species diversity Ecological Succession
How many different species? Who’s dominant? most abundant or highest biomass (total weight) Keystone species Ecological Succession
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Species diversity greater diversity = greater stability
Greater biodiversity offers: more food resources more habitats more resilience in face of environmental change
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The impact of reduced biodiversity
compare these communities suburban lawn agricultural “monoculture” “old field” In the early 1970's, a new corn genotype was released in the US; the "Texas male sterile" (TMS). Hybrids that contained this new characteristic had many desirable properties, and growers were excited about them, planting them over miles and miles of corn acreage in US. They were, of course, bred to be resistant to the most common corn diseases. However, they were not resistant to a previously unimportant strain of a fungal disease; the southern corn leaf blight (caused by the fungus Helminthosporium maydis). Ninety percent of the corn sowed in the US in 1970 contained the TMS trait and also shared genetic susceptibility to this pathogen. The fungus encountered all this acreage of susceptible host and wiped out one fourth of the US corn crop in 1970, a loss of over one billion dollars in production! If the corn acreage hadn't been such a monoculture, the fungus wouldn't have been able to spread as rapidly, as it would have encountered barriers of genetically resistant plants. Irish potato famine 1970 US corn crop failure
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Keystone species Influential ecological role
A species that has a large effect on its environment relative to its abundance A consumer organism (herbivore or carnivore) Low biomass in numbers
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Keystone Species diversity increases diversity decreases mussels out-compete other species Following the removal of Pisaster, richness fell from 15 to 8 species Pisaster promotes species diversity by preventing competitive exclusion by Mytilus
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Ecological succession
Sequence of community changes transition in species composition over time years or decades usually after a disturbance Mt. St. Helens
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Progress of Succession
Tolerance early species are weedy (r-selective) tolerant of harsh conditions Facilitation & Inhibition early species facilitate habitat changes change soil pH change soil fertility change light levels allows other species to out-compete
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Primary succession Begins with virtually lifeless area without soil, then… bacteria lichens & mosses grasses shrubs trees make soil {
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Secondary succession Existing community cleared, but base soil is still intact burning releases nutrients formerly locked up in the tissues of tree the disturbance starts the process of succession over again
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Climax forest Stable plant community dominated by trees
Representing final stage of natural succession for a specific location remains unchanged in species composition as long as site remains undisturbed birch, beech, maple, hemlock oak, hickory, pine
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Climax forest Hmm…Biomes?
taiga The species mix of a climax forest is dependent on the abiotic factors of the region solar energy levels temperature rainfall fertility & depth of soil temperate deciduous forest birch, beech, maple, hemlock
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