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Development of multicellularity
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Advantages of size Being big prevents others from eating you
Being big allows you to eat others Change in ratio of surface area to volume as size increases. Metabolic rate is somewhat size dependent
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Disadvantages of size Total energy requirements go up with increasing size In air and water, big causes more rapid sinking if not compensated for extensions of body or changes in specific gravity Diffusion becomes a problem
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There must be an overall advantage
Life as it evolved has become bigger (in bigger packages) Proximate and ultimate causes/effects
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How does it happen Divisions Multinucleation
Nuclear divisions--mitosis Cytoplasmic divisions--cytokinesis Multinucleation Syncitial--multinucleate, but no cell membranes. (fungi--saphrophytic/zoic Multicellularity--multiple mitoses and cytokineses
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Cellular adhesion Glue--cell cement Physical structures
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Advantages of multicellularity
Increase size Division of labor Specialized cells
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Disadvantages of multicellularity
Total volume of food goes up Diffusion distances increase making the physical process of diffusion less functional
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Volvocine method Volvox is a colonial algae Not much division of labor
Hollow sphere with cells in outside layer colony
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Protosponge approach Collar cells
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