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Chapter 24 Protists
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Kingdom Protista “Simple” eukaryotes Mostly unicellular
Some form colonies Some are coenocytic Multiple nuclei in one mass of cytoplasm A few are multicellular
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Most protists live in aquatic environments
Sixty major groups of protists Important in the biosphere Food for other organisms Photosynthetic protists supply oxygen
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Locomotion Modes of nutrition Pseudopodia Flagella Cilia
A few are nonmotile Modes of nutrition Autotrophs Heterotrophs
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Chlamydomonas
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Interactions with other organisms
Free-living Mutualism Commensalism Parasitism Habitats Most live in the ocean or freshwater ponds, lakes, and streams Parasites live in host’s bodily fluids
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Modes of reproduction Many protists reproduce both sexually and asexually Syngamy (union of gametes) Some protists are solely asexual
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Endosymbiont theory Certain organelles arose from symbiosis with prokaryotes Mitochondria arose from aerobic eubacteria Chloroplasts arose from cyanobacteria
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Primary and secondary endosymbiosis
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Classifying eukaryotes
Ultrastructure Fine details of cell structure Molecular data Ribosomal RNA Nuclear genes The protist kingdom is probably paraphyletic
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Eight monophyletic groups of eukaryotes
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Zooflagellates Mostly unicellular heterotrophs Move using flagella
Now separated into several monophyletic groups Excavates Discicristates
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Excavates live in oxygen-free environments
Diplomonads No mitochondria No Golgi complex One or two nuclei Up to eight flagella Giardia is a parasite
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Giardia
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Discicristates have disc-shaped mitochondrial cristae
Euglenoids About 1/3 are photosynthetic Inhabit freshwater ponds and puddles Trypanosoma causes African sleeping sickness
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Euglena gracilis
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Alveolates have flattened vesicles under the plasma membrane
Ciliates Move by hairlike cilia Micronuclei for sexual reproduction Macronuclei control metabolism Reproduce sexually by conjugation
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Paramecium, a ciliate
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Alveolates Dinoflagellates Apicomplexans
Mostly unicellular with two flagella Mostly photosynthetic Apicomplexans Parasitic Spore-forming Plasmodium causes malaria
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Dinoflagellates
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Heterokonts are motile with two different kind of flagella
Water molds Mycelium absorbs organic material Reproduce asexually with biflagellate zoospores Reproduce sexually with oospores Phytophthora causes plant diseases
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Mycelium around a dead insect
Saprolegnia
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Heterokonts Diatoms Golden algae
Mostly unicellular with shells containing silica Some diatoms are part of plankton Golden algae Mostly unicellular, biflagellate algae Both freshwater and marine
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Diatoms
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Heterokonts Brown algae Multicellular seaweed
Ecologically important in cooler ocean waters Kelps have leaflike blades, stemlike stipes, anchoring holdfasts, gas-filled bladders
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Brown algae
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Cercozoa are amoeboid cells that often have hard outer shells, called tests, through which cytoplasmic projections extend
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Cercozoa Foraminferans Actinopods Many-chambered tests with pores
Cytoplasmic extensions to move and obtain food Actinopods Mostly marine plankton Axopods to obtain food
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An actinipod
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Plants have chloroplasts bounded by inner and outer membranes
Land plants, red algae, and green algae are monophyletic Red algae and green algae are in kingdom Protista
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Red algae are mostly multicellular seaweeds
Ecologically important in warm tropical waters Red algae with calcium carbonate in their cell walls are important for reef building
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Red algae
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Green algae are diverse in size, structural complexity, and reproduction
May be the ancestors of land plants Multicellular forms do not have cells differentiated into tissues, unlike plants
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Green algae
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Amoebas move and obtain food using cytoplasmic extensions called pseudopodia
Capture and engulf food by surrounding it and forming a vacuole around it Entamoeba histolytica causes amoebic dysentery
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Plasmoidal slime mold Feeding stage is a multinucleate plasmodium that extends up to one foot in diameter In the reproductive stage, stalks called sporangia produce haploid spores
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Plasmoidal slime mold
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Cellular slime molds Feeding stage is individual amoeboid cells
During moisture or food shortage, they aggregate into a migrating pseudoplasmodium Forms stalked fruiting body containing spores
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Cellular slime mold
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