1. Lycopodiaceae

2. Clubmosses take their name
from the club-like strobilus
in which the sprorangia are
borne in most species.
In the clubmosses, each
sporangium is associated
with a leaf called a
sporophyll.

3. Leaves
Leaves = microphylls.

4. Microphylls can be four-ranked, as in Diphasiastrum digitatum ...

5. or they can be spirally arranged, as in Huperzia lucidula

6. Forked branching,
branches equal
Forked branching,
branches unequal

7. Branching in clubmossesp
Branching in clubmosses

8. Gemma of Huperzia lucidula
(and other species)

9. Sporangia are borne on the
upper, or adaxial, surface
of the sporophyll.

10. In some species, such as
shining clubmoss, sporangia
are borne along the stem
axis.

11. The sporangia themselves
are reniform (kidney-shaped)

13. Lycopodium clavatum, one of about 15 clubmoss spp. in Vermont

14. Huperzia Diphasiastrum Lycopodiella
Some of the clubmoss genera in Vermont
Lycopodiella

15. Lycopodium sitchense (alpine) an alpine clubmoss species
Diphasiastrum digitatum, a VT species
a tropical epiphytic species

16. Lubricant for draftsman’s work, surgical gloves, and condoms
Economic Uses
Lubricant for draftsman’s work, surgical gloves, and condoms
Flash Powder (obsolete)

17. Land Plant Phylogeny Bryophytes Lycophytes Ferns Gymnosperms
Angiosperms