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Journal of the Geological Society

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1 Journal of the Geological Society
The Spence Shale Lagerstätte: an important window into Cambrian biodiversity by Julien Kimmig, Luke C. Strotz, Sara R. Kimmig, Sven O. Egenhoff, and Bruce S. Lieberman Journal of the Geological Society Volume 176(4): July 4, 2019 © 2019 The Author(s). Published by The Geological Society of London

2 Locations and stratigraphy of the Spence Shale Lagerstätte.
Locations and stratigraphy of the Spence Shale Lagerstätte. (a) Map of the western USA showing the location of the Spence Shale. (b) Geological map (based on the USGS state maps for Google Earth Pro) of northern Utah and southern Idaho showing the principal localities within the Spence Shale. AC, Antimony Canyon; BF, Blacksmith Fork; CC, Cataract Canyon; CFC, Calls Fort Canyon; DC, Donation Canyon; EC, Emigration Canyon; HC, Hansen Canyon; HCR, High Creek; MH, Miners Hollow; ON, Oneida Narrows; PP, Promontory Point; SG, Spence Gulch; TMC, Two Mile Canyon. (c) Simplified stratigraphy of the Langston Formation. Julien Kimmig et al. Journal of the Geological Society 2019;176: © 2019 The Author(s). Published by The Geological Society of London

3 Sedimentology of the Spence Shale.
Sedimentology of the Spence Shale. (a) Lag deposit near base of ‘cycle 3’, consisting of biogenic debris, probably of echinoderm, brachiopod and trilobite remains. It should be noted how bedding bends around the millimetre-size echinoderm bioclasts. The matrix is carbonate-rich siliciclastic mudstone with varying amounts of sub-millimetre carbonate debris. (b) Carbonate-rich siliciclastic mudstone near top of ‘cycle 3’, with several millimetre-long black organic-rich flakes oriented parallel to bedding. Abundant silt-size carbonate particles in the matrix should be noted. Scale bar is 1 mm. Julien Kimmig et al. Journal of the Geological Society 2019;176: © 2019 The Author(s). Published by The Geological Society of London

4 Important private collectors.
Important private collectors. (a) Lloyd Gunther of Brigham City, UT. (b) Val (right) and Glade (left) Gunther of Brigham City, UT. (c) Paul Jamison of Logan, UT. Julien Kimmig et al. Journal of the Geological Society 2019;176: © 2019 The Author(s). Published by The Geological Society of London

5 Selected soft-bodied arthropods from the Spence Shale.
Selected soft-bodied arthropods from the Spence Shale. (a) KUMIP holotype of Meristosoma paradoxum from Miners Hollow, collected by the Gunther family. (b) KUMIP , Mollisonia symmetrica from Miners Hollow, collected by the Gunther family. (c) KUMIP , Waptia cf. W. fieldensis from Cataract Canyon, collected by Val and Glade Gunther. (d) KUMIP , Isoxys sp. from Miners Hollow, collected by Arvid Aase. (e) KUMIP , Tuzoia sp. with burrows under the carapace from Miners Hollow, collected by Phil Reese. (f) KUMIP , Leanchoilia superlata? from Miners Hollow, collected by Val and Glade Gunther. (g) KUMIP , hurdiid H-element from Miners Hollow, collected by the Gunther family. (h) KUMIP , Hurdia sp. appendage from Miners Hollow, collected by Paul Jamison. (i) KUMIP , arthropod appendage from Antimony Canyon, collected by Val Gunther. (j) KUMIP , Dioxycaris argenta from Miners Hollow, collected by the Gunther family (k) KUMIP , holotype of Yohoia utahana from Miners Hollow, collected by Paul Jamison. (l) KUMIP , holotype of Utahcaris orion from Antimony Canyon, collected by Ben Datillo. Scale bars represent 10 mm for (a), (e) and (l) and 5 mm for (b) (c), (d) and (f)–(k). Julien Kimmig et al. Journal of the Geological Society 2019;176: © 2019 The Author(s). Published by The Geological Society of London

6 Selected fossils from the Spence Shale.
Selected fossils from the Spence Shale. (a) KUMIP and KUMIP , Vauxia magna from Miners Hollow, collected by Rhiannon LaVine. (b) KUMIP , holotype of Wiwaxia herka from Miners Hollow, collected by Phil Reese and the Gunther family. (c) KUMIP , Wronascolex? ratcliffei from Miners Hollow, collected by Riley Smith. (d) KUMIP , Selkirkia spencei from the Wellsville Mountains, collected by the Gunther family. (e) KUMIP , Eldonia ludwigi from Antimony Canyon, collected by Lloyd and Val Gunther. (f) KUMIP , Sphenoecium wheelerensis from Miners Hollow, collected by the Gunther family. (g) KUMIP , Acinocricus stichus from Miners Hollow, collected by Paul Jamison. (h) KUMIP , Micromitra? sp. from High Creek, with chaetae preserved, collected by Paul Jamison. (i) KUMIP , ‘enrolled’ Amecephalus laticaudum from Miners Hollow, collected by Paul Jamison. (j) KUMIP , Zacanthoides liddelli from High Creek, collected by Paul Jamison (k) KUMIP , Oryctocephalus walcotti from Oneida Narrows, collected by the Gunther family. (l) KUMIP , holotype of Siphusauctum lloydguntheri from Antimony Canyon, collected by Lloyd Gunther. Scale bars represent 5 mm. Julien Kimmig et al. Journal of the Geological Society 2019;176: © 2019 The Author(s). Published by The Geological Society of London


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