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Cyperaceae The Sedge Family Juncaceae The Rush Family www. uvm
Cyperaceae The Sedge Family Juncaceae The Rush Family
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Character JUNCACEAE (Rushes) CYPERACEAE (Sedges) POACEAE (Grasses) Habitat Mostly wet areas Wetlands and upland woods Various Stems Terete in cross section Internodes solid with large pith Nodes not jointed Triangular (not always visible) Internodes usually solid Terete or ellipsoid Internodes hollow (to solid) Nodes jointed Leaves 3-ranked Blades flat to terete Open sheath Blades flat Closed sheath 2-ranked Inflorescence Basically cymose and often congested Each flower subtended by 2 bracts Spikelets Each flower subtended by one bract (scale) Each flower subtended by 2 bracts (lemma and palea Perianth Usually 6 chaffy tepals Absent or reduced to a number of bristles or scales Reduced to 2 (or 3) scales Fruit Loculicidal capsule Achene Caryopsis (grain)
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Cyperaceae The Sedge Family
Vegetative attributes: Mostly perennial herbs Stems often three-angled Nodes not swollen Internodes not hollow Leaves three-ranked, sheaths not split
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Dulichium arundinaceum, Three-way sedge
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Cyperaceae The Sedge Family
Reproductive attributes: Perianth, if present, of bristles or scales Androecium of three stames Ovary superior, 2-3 fused carpels Fruit an achene Each floret subtended by a scale Florets arranges in spikelets
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S Staminate florets Scale Spikelet Pistillate florets
Carex: no perianth, flowers unisexual
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Eriophorum, cotton grass:
Perianth of long bristles
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Cyperaceae: The Sedge Family 90 genera, 4000 species worldwide
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Vermont Genera: Carex (sedge) – 125 species
Scirpus (bulrush) – 17 species Cyperus (umbrella sedge, galingale) – 10 species Eleocharis (spike rush) – 9 species Dulichium (three-way sedge) – 1 species, common Eriophorum (cotton grass) – 5 species Rhynchospora (beak rush) – 4 species Cladium (twig rush) – 1 species, uncommon Fimbristylis (autumn fimbristylis) – 1 species, rare Bulbostylis (sand sedge) – 1 species
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Carex: perigynium
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A Key to Vermont Cyperaceae
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Perigynium? Yes: Carex No Flowers perfect and distichous Flowers spirally imbricate
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Distichous
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If perfect and distichous, it is either
Cyperus or Dulichium
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Spirally imbricate
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Spirally imbricate Tubercle?
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Yes, tubercle Spikelet one per culm Spikelets more than one per culm Eleocharis
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Tubercle minute Tubercle broad Bulbostylis Rhynchospora
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No tubercle Conspicuous bristles?
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Yes, bristles Very long: Eriophorum Shorter: Scirpus
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Bract of inflorecence one:
No conspicuous bristles Bract of inflorecence one: Scirpus
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Bracts of inflorecence multiple Flowers one per spikelet:
No conspicuous bristles Bracts of inflorecence multiple Flowers one per spikelet: Cladium
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Flowers several per spikelet
No conspicuous bristles, bracts of inflorescence multiple Flowers several per spikelet Leaves > 5 mm wide: Scirpus
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Leaves narrower Leaf sheaths long-ciliate Leaf sheaths short-ciliate Bulbostylis Fimbristylis
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