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PHYLUM MOLLUSCA (Molle= soft body)
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PHYLUM MOLLUSCA OVER 100,OOO LIVING SPECIES; MANY 1000’S OF FOSSIL SPECIES FOUND IN ALMOST ALL ENVIRONMENTS: MARINE, FRESHWATER, TERRESTRIAL INCLUDES SNAILS, SLUGS, CLAMS, SQUIDS, OCTOPUSES,
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CHARACTERISTICS Body Plan: three main parts- head, foot, visceral mass
body is unsegmented
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CHARACTERISTICS Mantle - specialized tissue, secretes shell, may be involved in feeding, reproduction and respiration Radula – for rasping; in all but one class Radula closeup
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ALL ORGAN SYSTEMS ARE PRESENT
Circulation - Open (Hemocoel) - Hemocyanin Respiratory System - gills (Ctenidia), lungs, mantle, epidermis Osmoregulation - Metanephridia also excretion
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ALL ORGAN SYSTEMS ARE PRESENT
Feeding - Radula (scraping) Bivalves - Filter feed Nervous System - Complex system of Ganglia
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ALL ORGAN SYSTEMS ARE PRESENT
Reproductive System - usually dioecious, monoecious in some Marine forms have free living trochophore larva Other larval stages may be present
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MOLLUSCAN CLASSES
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Class Monoplacophora “one plate”
all known from deep marine waters (>12,000 feet) have a single circular shell with radula
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Class Polyplacophora “many plates”- (chitons)
have 8 overlapping plates or shells reduced head, no eyes or tentacles all marine; usually live on rocky shores have radula, gills external fertilization scrape algae
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Polyplacophora Respiration - Gills in pallial groove
Circulation - Heart at posterior end - single Aorta Excretion - Nephridia Nervous System and sense organs - Ganglia and nerves Sub Radular organ - Feeding Esthetes - light sensing Osphradia - Sampling water
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Polyplacophora Reproduction - Sexes Separate
Usually external fertilization Trochophore larva
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Class Aplacophora “without plates”
Rare; lack shell; wormlike deep marine; burrowing species; predators About 320 species, not much is known of them Your book separates them into two separate classes: ! Caudofoveata ! Solenogastres
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Class Gastropoda Largest class of Molluscs
Most important feature is Torsion
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Torsion Twisting of body into loop Perhaps adaptation of veliger larva
Disadvantages: Reduction of organs on right side Excretion - Mouth moves near anus Adavntages Allows water to be drawn from the front
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Gastropod - Locomotion
Foot Ciliary action below the foot Operculum Swimming
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Gastropod - Nutrition Herbivores - Radula Carnivores Filter feeders
NERVOUS SYSTEM Similar to Chiton - but in the shape of “8”
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Class - Bivalvia Includes Calms, Oysters, Mussels
Has two Shells (Valves) Shell held together by Adductor muscle Head is lost Filter feeders Mantle secretes the shell
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MOLLUSCAN SHELL STRUCTURE
Periostracum; outer layer Prismatic; middle layer Nacre; inside layer- mother of pearl
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BIVALVIA Locomotion Circulation Excretion Foot (burrowers)
Active swimmers (Scallops) Circulation Open system Excretion Nephridia
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BIVALVIA Nervous System Reproduction 2 pairs of ganglia
Sense organs (statocysts, ocelli, ospharidia) Reproduction Sexes separate Internal fertilization Larva - Glochidium
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Class - Cephalopoda Includes Squid, octopus, Nautilus
Most advanced of Molluscs Well developed head and eyes Head has tentacles with suckers Squids are largest of invertebrates Locomotion by jet propulsion
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Class Scaphopoda (tusk shells)
slender, tubular shells open at both ends all marine have tentacles around mouth no defined head no gills, respiration across mantle
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