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PLANTS UBER ALLES (Plants as control freaks)
PLANT HORMONE PLANTS UBER ALLES (Plants as control freaks)
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OUTLINE History and Scientists involved
Control Systems in plants and animals Differences between plant and animal hormones Hormone action Plant Hormone and effects Experimental Evidence
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Scientists and History of Plant Hormones
Charles and Francis Darwin – initial proposition-1880 Boysen- Jensen – 1910 Went-1930 Skoog-1950 Kurosawa-1920 Addicott-US,Wareing-England Various fruit growers – 1960’s Australian Scientists
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CONTROL SYSTEMS Necessary for response to stimuli
Nervous system in animals Tropic response systems in plants and some invertebrates Hormonal systems in both plants and animals Endocrine Exocrine Hormonal mimics
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Differences between Plant and animal Hormones
Animal hormones produced by specific glands and/or cells. Plant hormones are produced by specific regions – ie. Meristems, cambriums ets. Most animal hormones are transported in some way via blood or by ducts Plant hormones generally must diffuse into adjacent cells or are carried by phloem
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Hormonal Action Most hormones act by way of the second messenger theory. This states that the hormone binds with a receptor in the cell membrane which then triggers an induction of chemical reactions causing specific protein to form
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HORMONAL ACTION Once induction has occurred, this frequently sets up a cascade effect which triggers other responses. Ie. Giberillins initiate water uptake in seeds which then causes a multitude of other responses such as production of amylase These effects are slow to act but long lasting in permanent changes.
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Plant Hormones Auxins Cytokinins Giberillins Abscissic Acid Ethylene
Florigen Artificial mimics Strigolactones
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Auxins Growth of apical meristem and supression of lateral meristem
Cellular eleongation in grass and herbs Differentiation of Vascular Tissue Fruit Development and parthenocarpy Wound repair and development of adventitious roots Calcium release – promotes activation of calmodulin – control of calcium receptors
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Cytokinins Stimulate cell division
Organogenesis in connection with auxins High cytokinin/auxin rations favor stem and leaf development Low rations favor root development Senescence and programmed death of cells –longer shelf life of vegetable in grocery stores
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Giberillins Rapid , Extensive growth of intact plants
Seed germination and removal of inhibitions Juvenile characteristics Increase of fruit size in seedless varieties Foolish seedling disease
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Ethylene Fruit Ripening –one rotten apple spoils the bushel
Flowering inhibited in most plants – exception Bromeliads Sex expression in diecious plants – holly Abscission – initiates break down of middle lamella at leaf /stem juncture
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Abscissic Acid Closes stoma in drought conditions
Creates dormancy in seeds and buds Formation of abscissic layer in leaves causing death and drop of leaf Counteracts stimulatory effects of other hormones
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Florigen Stimulates flowering
Has been proven to exist but has never been isolated Experiments with long day and short day plants
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Strigolactones Stimulates mycorrhizial fungal attachment to roots
Stimulates formation of lateral buds(Fibionacci numbers) Helps seeds break dormancy after fire (Australian Fire Ecology)
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