1. Soil organic matter Soil organic matter soil bio- geochemistry population dynamics & disturbance plant biogeography primary production & growth Vegetation climateCO 2 2

3. Vegetation representation in LPJ-GUESS

5. Cohort nitrogen uptake capacity = f (C root,N:C plant ) Soil retranslocation N demand = f (V max,∆C,...) Vegetation N deposition N fixation = f (ET) N leaching N trace gases N leaf N root N sap N store metab surface microbial struct FWD CWD metabstruct surface humus soil microbial slow SOM passive SOM Leaf & wood litterroot litter N avail Litter C,N