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The Burgess Shale A Window on the Cambrian Explosion
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Konservat Lägerstatte Konservat – preservation Lägerstatte – “treasure house” Konservat Lägerstatte – a location or stratigraphic unit with unique and remarkable preservation of fossils Fossils may be uncommonly abundant or rare Fossils are typically articulated Soft tissues may also be preserved
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Burgess Shale Middle Cambrian Shale – deep, dysoxic Yoho National Park Canadian Rocky Mountains The world’s most important Lägerstatte?
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The Cambrian Explosion Widespread appearance of abundant fossils Evolution – Explosive Adaptive Radiation A Cambrian “arms race”? or Creation of Habitat – Passive Margins of Rifted Rodinia OR… Taphonomic Event - Made possible by development of shells Diverse taxa develop ability to make shells at once Chemical threshold in oceans? Atmosphere? OR… ALL OF THE ABOVE????
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The Burgess Lägerstatte Excellent preservation Preservation favored by Fine grained sediment Anoxic to dysoxic environment Obrution Events – rapid burial of live organisms Unique Organisms – some have no modern descendents Ancestors of many extant taxa “Wonderful Life” – variations and contingency
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Anomalocaris
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Marella (“lace crab”) and Wiwaxia (an armored “tribble”?)
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Opabinia, Yohoia, Sidneyia
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Halucigenia First Reconstruction Recent Reconstruction
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Canadia (a worm) and Pikaia (first notochord) Canadia Pikaia
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Chenjiang Biota – Maotianshan Shale Early Cambrian of China Preservation of soft- bodied biota Similar to Burgess Shale But 15 MY older!
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