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Competition
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Species can compete for two resources and can coexist when 2 conditions are met:
1) The habitat must be such that one species is more limited by one resource and the other species is more limited by the other resource. 2) Each species must consume more of the resource that more limits its own growth.
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Another look at plant strategies – from Peter Grime
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Grime’s C-S-R strategies
Competitive - plants such as perennial herbs, shrubs and trees typically have dense leaf canopies, high growth rates, low seed production and relatively long life spans Stressful -stress-tolerant plants that often have small evergreen leaves, low growth rates, low seed production and long life spans Disturbed environments - the plants are primarily Ruderals – plants which are usually small, grow rapidly, short life spans and produce many seeds
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Grime’s C-S-R strategies
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Fynbos – South African Mediterranean shrubland
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Fynbos burning
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Fynbos recovery
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Competition in Nature – the Birth of Experimental Ecology
Joe Connell
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Acorn barnacles growing on an old car tire
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Acorn barnacles on the rocky coast
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Sceloporus – Fence lizard
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Urosaurus – long tailed brush lizard
aka tree lizard
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Competitive Release Competitive Release is a prediction from examining the competitive exclusion principle that in the absence of competition a species should expand its niche
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Chalcophaps indica – Emerald Dove
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Chalcophaps stephani – Stephen’s Dove
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Gallicolumba rufigula
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Character Displacement
Character displacement is a measurable physical difference between two species which has arisen by natural selection as a result of the selection pressure on one or both to avoid competition with each other - here we assume that environment is the same at all locations
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Hydrobia ulvae Hydrobia ventrosa
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Hydrobia ulvae – note size
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The Ghost of Competition Past
Joe Connell
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Patterns like character displacement or competitive release can be caused by several things:
the pressure of current competition causes the pattern competition which occurred in the past may have driven natural selection to cause the pattern we see today - "the ghost of competition past" competition in the past eliminated a number of other species, leaving behind only those that were different in the use of habitat the species may have evolved independently and in different ways and have never competed with each other the species may differ in their niches, but not enough to coexist in a stable environment, however the environment varies and thus prevents competition from reaching its predicted end result
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Blackburnian warbler Cape May warbler Black-throated green warbler Bay-breasted warbler Yellow-rumped
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Ghost of Competition Past in Israeli Rodents
Gerbillus allenbyi Meriones tristami
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