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ניורוביולוגיה ומדעי המח חלק 2 – מערכת הראייה Introduction to Neurobiology Part 2 – The Visual System Shaul Hochstein
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ניורוביולוגיה ומדעי המח Introduction to Neurobiology Introduction to Sensory Systems The Visual Cortex “From Neuron to Brain”: Chapters 20 & 21: pages 411-433; Signaling in the LGN & primary visual cortex & Functional architecture of visual cortex
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ניורוביולוגיה ומדעי המח Introduction to Neurobiology Introduction to Sensory Systems What do we mean by “why” in science? What is the primary goal of neuroscience?
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4 principles of physiological research
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Study of receptive fields is equivalent to study of computation; (from input/output can deduce computation). Study of RF linearity/nonlinearity essential for deriving mechanisms. Division into neuron classes basic for visual system, differentiating processing pathways, using one, the other, or a combination of streams. Analysis of response types may aid in understanding neuron role/function.
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ניורוביולוגיה ומדעי המח Introduction to Neurobiology Introduction to Sensory Systems To where are retinal signals transmitted?
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David Hubel and Torsten Wiesel
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ניורוביולוגיה ומדעי המח Introduction to Neurobiology Introduction to Sensory Systems Mapping the receptive field of visual system neurons using small spots of light or dark. Very effective in RGC & LGN. Very problematic for Visual Cortex.
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This is nonsense: The responses of cortical Simple cells depend largely, essentially and dramatically on intra-cortical networks.
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This gives a false impression: We do not know that Complex cells depend on Simple cell input and it is clear that end-stopping exists in Simple cells and is not a “hyper- complex” property.
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The primary visual cortex maintains: 1- visual hemifield location map 2- size discrimination (small and large receptive fields) 3- color coding (in monkey; in “blobs”) The primary visual cortex begins selectivity to: 1- orientation 2- ocular dominance and binocularity, leading to 3- stereo depth perception 4- receptive field complexity (Simple vs. Complex cells) 5- end-stopping 6- motion direction sensitivity
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Visual field magnification factor
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