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Animal adaptations Outline: Acquisition of energy and nutrients Respiration Homeostasis Water balance Biological rhythms Readings: Chapter 7
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Energy and nutrient acquisition
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Detritivores
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Herbivores
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Types of herbivores Grazers - leaf tissue Browsers - woody tissue Granivores - seeds Frugivores - fruit Nectivores - nectar Phloem feeders - sap High cellulose (fiber), low protein Animals can’t digest cellulose (no cellulase enzymes) Need symbiotic bacteria, protozoa
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Ruminants (e.g. cows, sheep, deer)
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Non-ruminants (e.g. rabbits, horses)
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Coprophagy = ingestion of feces E.g. Lagomorphs (rabbits, hares & pikas) E.g. Detritivores
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N and food quality For herbivores, food quality increases with increasing N content In animals, C:N ~ 10:1 In plants, C:N ~ 40:1 herbivores limited by N availability –Highest in growing stems, leaves, buds –Decreases as plant ages Herbivores usually born in spring
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Carnivores
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Composition of food similar to own tissues --> simple stomach --> small caecum Need to get enough food
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Omnivores Feed on > 1 trophic level, e.g. plants and herbivores Diet varies with season, life cycle
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Diet breadth 1.Generalists: “polyphagous” – eat >1 prey species 2.Specialists: “monophagous” – eat one prey species – or eat specific part of prey E.g. seed-eating birds Specialists are usually Short-lived (active only when food is available) Highly adapted to a specific food type (can’t use any other)
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C 6 H 12 O 6 + 6O 2 6CO 2 + 6H 2 O + ATP RESPIRATION
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HOMEOSTASIS
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THERMOREGULATION
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TEMPERATURE REGULATION TYPE OF HEAT PRODUCTION: Endothermy: - heat from within Ectothermy: - heat from without Heterothermy - employ endo and ectothermy in different situations TEMPERATURE VARIATION: Homeothermy - constant temperature Poikilothermy - variable temperature
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TEMPERATURE REGULATION poikilotherms
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Operative temperature range
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TEMPERATURE REGULATION poikilotherms Acclimatization
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Lizards and snakes: body temperature varies only 4-5 o C/day TEMPERATURE REGULATION poikilotherms
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TEMPERATURE REGULATION homeotherms
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Endothermy – ectothermy tradeoffs
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Endothermy tradeoff
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Because of their small size (high surface: volume ratio) and their need to invest energy in growth, juvenile birds and mammals are often ectothermic, obtaining heat from their parents. Conserving energy – ectothermy for juveniles
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Bears are not true hibernators; their body temperature drops only a few degrees, and they are relatively easily awakened Conserving energy – hibernation
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Conserving energy – countercurrent heat exchange without with
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Releasing energy – countercurrent heat exchange RETE
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Adaptations to aridity and heat
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Water balance in aquatic environments Freshwater organisms: hyperosmotic (water wants to move inside of organism Marine organisms: hypoosmotic (water wants to move outside of organism
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Controls on activity
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Human diurnal cycle
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Life history strategies Outline: Types of reproduction Mating systems Sexual selection Energy and timing of reproduction Offspring Habitat selection Environmental influences Readings: Ch. 8
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A simple life history Life history = schedule of birth, growth, reproduction & death
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Types of reproduction Asexual or sexual Different forms of sexual reproduction
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Simultaneous hermaphrodites
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Sex change
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Mating system Strength of bond: – Monogamy (strong) - Promiscuity (no bond) Types of bonds: – Monogamy (one-to-one) – Polygamy (one-to-many) Polygyny (one male, many females) Polyandry (one female, many males)
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POLYANDRY: African Jacana
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Sexual selection Intrasexual selection – male-to-male or female-to-female competition for the opportunity to mate
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Sexual selection Intersexual selection – differential attractiveness of individuals
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Reproduction is costly
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Timing of reproduction Semelparity - reproduce once and die Iteroparous - reproduce throughout lifetime
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European grasshopper, Chorthippus brunneus An iteroparous summer annual
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Pigweed, Chenopodium album A semelparous summer annual
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Semelparous perennials Coho salmon: a long-lived semelparous animal Dies after spawning (2-5 yrs) Overlapping generations
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Bamboo Both genets and ramets are semelparous. Genets can live for 200 years before the simultaneous flowering of all ramets. Semelparous perennials
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Parental investment
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Fecundity
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Reproductive tradeoffs
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r and K strategists
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For next lecture: Please read Chapter 9, 10, 11, 12 65
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