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Evolution
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Fixed Species Concept u The creator had designed each and every species for a particular purpose.
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Result u No evolution. u Created the viewpoint that all species could be identified and named (Taxonomy). A major factor in the Linnaeus classification system.
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Theory u Fossils were the remains of species lost due to catastrophe. u No new species originated; species could only be lost over time. u Result - No evolution.
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James Hutton u 1795 - Gradualism u Profound change is the cumulative product of slow, but continuous processes.
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Result u Changes on the earth were gradual, not catastrophic.
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Charles Lyell u 1797 - 1875. u Incorporated Hutton’s gradualism into a theory called Uniformitarianism.
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Uniformitarianism u Geological processes have operated at the same rate over the Earth’s history.
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Result u The Earth must be VERY old. (much older than 6000 years of the fixed species concept). u Idea that slow and subtle processes can cause substantial change.
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Jean Baptiste Lamarck u Published theory in 1809. u Theory - Life changed from simple to complex over time.
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Lamarck u Fossils were the remains of past life forms. u Evolution did occur.
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Lamarck’s Mechanisms 1. Use and Disuse - u Body parts used to survive become larger and stronger. u Body parts not used to survive deteriorate.
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Lamarck’s Mechanisms 2. Acquired Characteristics u Modifications acquired by use/disuse were passed on to offspring.
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Lamarck’s Mechanisms 3. Natural Transformation of Species ….species changed with every generation… extinction never occurred, organisms simply changed overtime.
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Problems with Lamarck’s Theory u No knowledge of genetics. u Acquired traits are not transmitted offspring.
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To Lamarck’s Credits u Did suggest correctly the role of fossils in evolution. u Did suggest that adaptation to the environment is a primary product of evolution.
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Charles Darwin u Father of the modern theory of evolution. u Theory - Descent with Modification.
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Darwin's Background u Trained as a Naturalist (after trying religion and medicine).
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Voyage of the Beagle
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Result u Darwin's training and travel opportunities allowed him to formulate and support his ideas on Natural Selection.
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Galapagos Finches
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Galapagos Islands
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Unique animals Marine IguanaTortise
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Alfred Wallace - 1858 u Paper on Natural Selection identical to Darwin's ideas.
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Result - July 1, 1858 u Dual presentation of the Wallace-Darwin ideas to the Linnaean Society of London.
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Darwin - 1859 u Publication of "The Origin of Species”
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Comment u Darwin best remembered for the theory because of his overwhelming evidence and because he published.
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Darwinian View u History of life is like a tree with branches over time from a common source. u Current diversity of life is caused by the forks from common ancestors.
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Example
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“The Origin of Species” u Documented the occurrence of evolution. u Suggested that the mechanism for evolution was Natural Selection.
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The Facts: Fact 1 - All species reproduce themselves exponentially.
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Fact 2 - Most populations are normally stable in size. Fact 3 - Natural Resources are limited (finite).
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Inference 1 u The large number of offspring must compete for the finite resources. u Result - Most offspring die.
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More Facts Fact 4 - No two individuals in a population are exactly alike. Fact 5 - Variation is inheritable.
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Inference 2 u Those individuals whose inherited characteristics fit them best to their environment survive and reproduce.
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Inference 3 u Offspring inherit the favorable characteristics. Populations shift over time as the favorable characteristics accumulate.
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Nature u Determines which characteristics are favorable. u Determines who survives. u Result - “Natural Selection”
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Natural Selection in action
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Artificial Selection u When man determines the characteristics that survive and reproduce. u Result - the various breeds of animals and plants we’ve developed.
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Ex - Mustard Plant OriginalCultivars
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Evolution Success Measured By u Survival u Reproduction u Whoever lives long enough and has kids is the “winner” in evolution.
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Requirements u In order for Natural Selection to work, you must have: u Long periods of time. u Variations within a population.
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Subtleties of Natural Selection 1. Populations are the units of Evolution. 2. Only inherited characteristics can evolve.
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Comment u Acquired characteristics may allow a species to evolve "outside" of Natural Selection. u Ex: culture, learning
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Evidences of Evolution 1. Biogeography 2. Fossils 3. Taxonomy 4. Comparative Anatomy 5. Comparative Embryology 6. Molecular Biology
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Biogeography u The geographical distribution of species. u Problem: u Species mixtures on islands u Marsupials in Australia
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Evolution Viewpoint u Biogeographical patterns reflect descent from the ancestors that colonized that area.
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Fossils u Relics or impressions of organisms from the past. u Problem: u Show changes over time from simple to complex. u Many fossils don't have descendants.
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Evolution Viewpoint u Life has changed over time. u Many species failed to survive and became extinct.
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Comments 1. Fossilization is a rare event. 2. Only hard parts fossilize well. 3. Problem in finding fossils. 4. Interpretation. 5. Missing Links.
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Taxonomy u Science of Classification.
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Main Categories u Kingdom u Phylum or Division u Class u Order u Family u Genus u Species
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Comparative Anatomy Problems 1. Homologous Structures - Common "building plan” with divergent functions. Mammal forelimbs
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Comparative Anatomy Problems 2. Vestigial Organs - Rudimentary structures of marginal, if any, use. Whale Legs
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Human Example
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Evolution Viewpoint u Remodeling of ancestral structures as their functions or adaptations changed.
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Comparative Embryology u Problem - closely related organisms go through similar stages in their embryonic development. u Ex: Gill pouches in vertebrates
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Evolution Viewpoint u Ontogeny is a replay of Phylogeny. (Development reflects descent)
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Molecular Biology u Study of Evolution at the DNA or protein levels. u Problem - related species have similar DNA sequences.
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Evolution Viewpoint u Related species share a common ancestrial DNA. The closer the relationship, the more similar the DNA sequences should be.
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