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Annelids – Defining Characteristics
Segmented worms Coelomates Trochophore larvae protostome
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Segmentation
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Annelids – Feeding Filter feeders – use gills to capture food particles from water Others use pharynx to either suck in food or to extend out and get it. Some annelids have jaws on their pharnyx Christmas tree worm
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Nereis – (clam worm) -has jaws for eating and fighting
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Respiration Aquatic annelids use gills
Earthworm cuticle Aquatic annelids use gills Terrestrial annelids exchange gases through their skin – their skin must remain moist for this purpose Cuticle on the earthworm helps prevent desiccation Feather-duster worm – gills exposed (red) - tentacles for gathering food
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Internal Transport Closed circulatory system
Dorsal surface has blood moving toward the head, ventral surface moves blood away from the head Each segment has a ring vessel connecting the ventral with the dorsal Large ring vessels at anterior end act as a “heart” to pump the blood If no heart is present, the bodies muscular contractions move the blood around
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Excretion Solid wastes out the anus
Metabolic (liquid) wastes are eliminated by nephridia (like the nephrons in our kidneys) 2 nephridia/segment
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Response Brain on dorsal surface
Nerves travel from the brain, around the gut and a main nerve cord travels down the ventral surface Brain = #11 Ganglia = #12 Ventral Nerve cord = #13
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Sense organs Best developed in free living annelids
Most of these have sensory tentacles, statocysts, chemoreceptors, and “eyes” Some form images, some are light-detectors Some have sensory organs that detect vibrations (like our tympanic membranes) Spaghetti worm underground With tentacles exposed
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Self Defence Burrow into ground or swim away
Setae (bristles) anchor worms in the soil Marine fan worms make tubes of CaCO3 to hide in Tube worms
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Self-Defence Marine fireworms have poisonous bristles (burns)
Carnivorous annelids have jaws to bite their predators
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Movement longitudinal muscles shorten the body
Circular muscles contract to decrease the diameter of the worm Coordination of these sets of muscles allow the worms to wiggle, burrow and swim
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Reproduction Some reproduce by budding
Some have separate sexes and have external fertilization Palolo worms congregate at the surface and release sperm or eggs at the same time Feast for the predators (humans included!)
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Earthworms are hermaphrodites and exchange sperm
Sperm is stored in sacs until eggs are ready Mucus ring is secreted by the clitellum – sperm and eggs are deposited there and the ring slips off the worm, forming a cocoon for the fertilized eggs
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Common Characteristics in all Annelids
Unidirectional Digestive System Closed Circulatory System First sign of a ‘heart-like’ structure Nephridia used for excretion Primitive Brains present
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Class Polychaeta – bristleworms
Mainly marine worm Have padddle-like appendages with bristles Show cephalization
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More Polychaetes
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Polychaetes
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Class Oligochaeta - Earthworms and Relatives
Earthworms + Freshwater worms Aquatic – eg tubifex (red, thread-like worms sold for fish food) Oligo – means few (as in few bristles) Castings are excellent fertilizer
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Class Hirudinea Commonly known as the LEECHES
Fresh and saltwater species (~300) Few land species Muscular proboscis is forced into skin Others slice skin with their jaws
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Leeches Hirudin – anti-coagulating chemical is used to keep host’s blood flowing Some also use an aesthetic so the host does not know they are there Used for medicinal purposes
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Annelids aerate the soil by burrowing
Return nutrients to the soil completing decomposition of material Many annelids are the basis of food chains
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Internal Anatomy
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Identifying practice:
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