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Community ecology Outline: Community structure: attributes Factors influencing the structure of communities Community dynamics Chapter 16-18
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Community attributes # of species Relative abundance of species Nature of species interactions (food webs) Physical structure
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Community structure Species richness (# of species within community) 10
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Community structure Relative abundance (% each species contributes to the total number of individuals)
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Stand one Stand two
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Simpson’s diversity index D = 0 - 1 –0: high diversity –1: low diversity Stand one (Table 16.1): D=0.13 Stand two (Table 16.2): D=0.36
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Dominance Yellow-poplar
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Food webs
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Keystone species
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Functional groups Feeding level Exploitation of common resources (guilds) Photosynthetic pathway Shade tolerance Life history
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Physical structure
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Zonation
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supratidal intertidal subtidal
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Association Relatively consistent species composition Uniform general appearance Distribution that is characteristic of a particular habitat
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Organismic concept of communities
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Continuum concept of communities
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Factors controlling community structure
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Fundamental niche
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Species interactions
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Keystone predation
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Apparent competition
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Indirect commensalism
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Top-down vs. bottom-up control
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The number of trophic levels regulates plant, herbivore and carnivore numbers B/comp: numbers limited by availability of resources (bottom-up regulation). Competition strong, predation weak. T/pred: numbers limited by predation (top-down regulation). Competition weak, predation strong. # trophic levels:
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Stress tolerance and competition
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Smooth cordgrass Salt meadow cordgrass Black needle rush
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Environmental heterogeneity
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Relationship between the number of plants per 300m 2 plot beside the hood river, NWT, and an index (ranging from 0 to 1) of spatial heterogeneity in abiotic factors associated with topography and soil. More spatially heterogeneous plots had higher species richness.
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Environmental quality
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Plant species diversity in a control plot and a fertilized plot in the Parkgrass experiment in Rothamstead, England. Fertilized plots have lower species diversity. The Parkgrass experiment, which began in 1856, is the longest running ecological experiment.
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Community stability Types of stability: 1.Resilient community: returns to former state after disturbance 2.Resistant community: changes little in response to disturbance
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Community dynamics: Succession
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Pioneer species Late successional species Primary succession
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Secondary succession
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beach grassshrubspinesoak
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Primary succession: newly exposed substrate
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Dryas sp. An early succession species on glacial moraines in Glacier bay. Dryas is a symbiotic N-fixing plant
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Populus trichocarpa Salix arctica After Dryas, cottonwood and willows become established
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Alnus incana Alders become the dominant tree after 50 years
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Tsuga mertensianaPicea sitchensis “Climax”: mixed spruce-hemlock forest
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Secondary succession: after disturbance
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Density Autogenic vs. allogenic change
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Allogenic environmental change
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Oak-pine forest Species diversity during succession
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Intermediate disturbance hypothesis
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HighLow
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In New Zealand streams, less disturbed sites support more complex communities (i.e., communities with larger, more connected food webs)
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Bark beetle/wood- boring beetle Fungi/bacteria Predatory insects Moss and lichen Invertebrates/ mice/ salamanders Succession of heterotrophs
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Changes over geologic time
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Concept of community revisited
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