1. Chapter 40 The Senses

2. I.Sensory Reception A. Sense organs contains specialized sensory receptor cells that detect stimuli & send the information to CNS

3. B. Signal Transduction – mechanism of changing environmental stimuli into neural impulses (action potentials) 1. Occurs in plasma membrane of receptor cell. Stimulus opens & closes membrane channels (Na + ) 2. Produces a receptor potential, or generator potential, which is a change in membrane polarization

4. 3. Most receptor cells are producing receptor potentials all the time, but at a slow steady rate, even when there is no stimulus a) When a stimulus IS present the receptor cell releases more transmitter than usual & increases the rate of action potentials generated in sensory neuron. b) This signals the brain that a stimulus was detected.

5. C. How do identical action potentials communicate different information? 1. By WHICH interneurons the sensory neurons are synapsing with inside of the brain. D. How do action potentials communicate information about intensity of stimuli? 1. By RATE of transmission of action potentials to the brain  Stronger stimuli generate more frequent action potentials being sent to the brain

6. E. Sensory Adaptation 1. The tendency of sensory receptor cells to become less sensitive when they are stimulated repeatedly. a) This will trigger fewer action potentials b) Brain may lose awareness of the stimuli

7. II. Specialized Sensory Receptors A. Skin contains: 1. Pain receptors 2. Thermoreceptors (heat & cold) 3. Mechanoreceptors (touch) a) In each of these sensors one neuron serves as the receptor and the sensory neuron. b) Dendrites transduce stimuli & axons send info. to CNS

8. B. Pain receptors 1. Naked receptors 2. Serve to warn us of danger or injury 3. Located everywhere except brain C. Thermoreceptors 1. Skin receptors detect heat & cold 2. Deeper sensors monitor temperature of blood a) Hypothalamus is body’s thermostat

9. D. Mechanoreceptors 1. Very diverse 2. Stimulated by touch, pressure, stretching, motion and sound 3. All work by some force bending or stretching membrane of receptor cell. 4. As membrane changes shape, it becomes more permeable to + ions a) Touch receptors around bases of hairs detect hair movements b) Stretch receptors monitor positions of body parts

10. E. Hair Cells 1. Detect sound waves & other movements in air or water 2. “Hairs” are specialized cilia or microvilli. a) When hair is bent in one direction, membrane is stretched, increasing permeability to + ions. Increases action potentials. b) When bent in opposite direction, less transmitter is released & action potentials decrease.

11. F. Chemoreceptors 1. Located in nose and taste buds 2. React to chemicals in the external or internal environment a) CO 2 detectors in blood b) Silkworm moth antennae detect sex attractant produced by female.

12. G. Electromagnetic receptors 1. Respond to electricity, magnetism and light a) Some fish detect electric fields given off by their prey. b) Electric fish discharge electrical bursts into the water to stun prey c) Many species have magnetite, an iron-containing mineral May help them navigate in world

13. 2. Photoreceptors a) Most common type of electro- magnetic receptor b) Detect light (visible, infrared, or ultraviolet) c) Snakes have two kinds: Normal eyes for visible light Infrared detectors - located in pits below normal eyes. Detect body heat from up to 1 m away. d) Lizards may have UV detectors in parietal eye

14. Vomeronasal OrganParietal Eye

15. III. Taste & Smell A. Receptors for both senses are chemoreceptors. 1. They interact with each other B. Smell 1. Receptors located in noses of vertebrates. Contain cilia. 2. Located on antennae of many insects

16. Olfactory Cell Location & Anatomy

17. C. Taste 1. In humans, receptors are located in taste buds on tongue. a. Four types of taste buds:  Sweet  Sour  Salty  Bitter 2. Insects have chemoreceptors for taste in sensory hairs on their feet

18. Taste Buds in Humans

19. IV. Vision A. Three types of photoreceptors: 1. Eye cup (eye spot) - Planaria 2. Compound eye - arthropods 3. Single-lens eye - vertebrates

20. B. Eye Cup 1. Photoreceptor cells contain molecules of a visual pigment a) These molecules absorb light b) Cause light energy to be transduced c) Action potentials sent to brain d) Do NOT form images. Show intensity and direction of light

21. C. Compound Eye 1. Have lenses that focus light and form images a) Consist of tiny light-detecting units called ommatidia b) Each has its own transparent covering called a cornea & a lens c) Lens focuses light on several photoreceptor cells d) Function as acute motion detectors e) Some insects see UV radiation

22. Compound Eye

23. Nectar Guides

24. D. Single-lens eyes 1. Anatomy - know functions of each a) Sclera b) Choroid c) Cornea d) Conjunctiva e) Iris f) Pupil g) Lens h) Retina i) Fovea j) Blind spot k) Vitreous humor l) Aqueous humor

25. Blind spot

26. 2. How to Focus (Accommodation) a) Lens focuses light onto retina. Bends rays b) Two ways to focus: Rigid lens moves back & forth Muscles change shape of an elastic lens  Thicker lens - bends light more sharply

27. c) Focusing in Humans Shape of lens controlled by ciliary muscles attached to choroid  When eye focuses on nearby object, these muscles contract (get a smaller diameter)  This makes ligaments that hold lens become slack.  Elastic lens then becomes thicker and rounder Reverse happens when focusing on distant object. Ciliary muscle relaxes. Lens becomes flatter.

28. Focusing of the Human Eye

29. d) Know how corrective lens work to correct each of the following conditions:  Near-sightedness (Myopia) Eyeball longer than normal Can’t focus well on distant objects  Far-sightedness (Hyperopia) Eyeball shorter than normal Can’t focus well on nearby objects  Astigmatism (misshapen lens or cornea)

30. 3. Photoreceptors at Work a) About 130 million photoreceptors of two types: Cones - Color receptors. Provide very sharp images but need plenty of light. Don’t function at night Rods - very sensitive to light. Allow us to see in dark situations but only in shades of gray.

31. b) Arrangement of Photoreceptors:  Cones - Densest in the fovea. Best to look directly at object. Allows for sharp daytime vision  Rods - Absent from fovea. Greatest density at outer edges of retina. Best to look at an angle at night to see dim objects more clearly

32. Photoreceptors of the Human Eye

33. c) Visual Pigments of Photoreceptors:  Each rod & cone contains a stack of disks containing light-absorbing pigments.  Rod pigment = rhodopsin  Cone pigment = 3 different photopsins Have 3 types of cones: blue green red See spectrum picture on next slide

34. Absorption Spectrum of Photoreceptors

35. d) Visual Sensory Transduction:  When pigments absorb light, they change chemically  Changes permeability of membrane  Creates receptor potential which will ultimately be sent to the brain  Complex integration occurs due to arrangement of cells in the retina

36. e) Visual Signal Pathway:  Receptor tips embedded in BACK of retina  Light must pass through the several other layers of cells which are between receptors and the incoming light  Information transduced by rods & cones must then pass in the opposite direction through the network of neurons  Axons make up optic nerve to brain

37. Structure & Function of the Retina

38. V. Hearing A. Involves hair cells in fluid-filled canals. B. Ear is composed of 3 regions: 1. Outer ear consists of pinna and auditory canal a) Collects sound waves b) Directs sound to the tympanic membrane (eardrum) Eardrum

39. 2. Middle Ear a) Eardrum vibrates at the same frequency as the original sound. Frequency is measured in hertz (Hz) = # vibrations/sec. b) Eardrum causes middle ear bones (malleus, incus, stapes) to vibrate in sequence. c) Bones transmit vibration to the oval window, which is much smaller in diameter than eardrum This amplifies the vibrations

40. Anatomy of the Human Ear

41. d) Middle ear is connected to pharynx via the Eustachian tube  Allows air pressure to be equalized on each side of the eardrum

42. 3. Inner Ear a) Oval window transmits sound waves to the snail-shaped cochlea b) Cochlea is a long-coiled tube that contains 3 separate fluid-filled canals:  Vestibular - upper canal. Begins at oval window.  Tympanic - lower canal. Connects to vestibular canal & ends at round window  Cochlear - between the other two

43. Cochlea

45. c) Organ of Corti  Actual organ of hearing  Long, thin spiral within cochlear canal  Consists of hair cells embedded in the basilar membrane  Tips of hairs touch overlying shelf Basilar membrane

46. Mechanoreceptors for Hearing

47. C. Physiology of Hearing 1. Oval window causes waves in the fluid of the vestibular canal 2. At top of coil, pressure waves enter the tympanic canal 3. Waves start the cochlear canal and basilar membrane to vibrating 4. Causes some hair cells to be bent.  Ion channels open and K + enters 5. Receptor potential is generated. 6. Action potential sent down sensory neuron axon (auditory nerve) to brain

48. D. How are Volume & Pitch Determined? 1. Volume (measured in decibels)  Louder sounds produce bigger waves which create more bending of hair cells  This sends more action potentials down auditory nerve towards brain  Human hearing ranges from 0 dB to 120 dB (point of intolerable pain)  > 90 dB sounds can destroy hairs

49. 2. How is Pitch (Frequency) Detected? a) Basilar membrane is not uniform in width along its spiraling length  End near oval window is narrower  End near tip of coil is wider b) High-pitched sounds generate high frequency pressure waves Causes narrower parts of basilar membrane to vibrate c) Low-pitched sounds cause the wider parts of basilar membrane to vibrate * Humans hear from 20-20,000 Hz

50. Frequency Detection

51. VI. Balance A. Involves two types of receptors: 1. Semicircular canals 2. Two chambers(saccule & utricle) B. Canals - detect rotational motion of head in any of 3 planes C. Chambers - detect position of head in relation to gravity

52. Mechanoreceptors for Equilibrium

53. Sensory Receptors in Other Animals