1. Chapter 13 Premodern Humans

2. Chapter Outline When, Where and What Premodern Humans of the Middle Pleistocene A Review of Middle Pleistocene Evolution Middle Pleistocene Culture

3. Chapter Outline Neandertals: Premodern Humans of the Late Pleistocene Culture of Neandertals Genetic Evidence Trends in Human Evolution: Understanding Premodern Humans Issue: The Evolution of Language

4. The Pleistocene The Pleistocene, often called the Ice Age, was marked by advances and retreats of massive continental glaciations.  At least 15 major and 50 minor glacial advances have been documented in Europe.  Hominids were impacted as the climate, flora, and animal life shifted. Middle Pleistocene (780,000– 25,000 y.a.) Upper Pleistocene (125,000–10,000 y.a.)

5. Interglacials Climatic intervals when continental ice sheets are retreating, eventually becoming much reduced in size. Interglacials in northern latitudes are associated with warmer temperatures, while in southern latitudes the climate becomes wetter.

6. Glaciations Climatic intervals when continental ice sheets cover much of the northern continents. Glaciations are associated with colder temperatures in northern latitudes and more arid conditions in southern latitudes, most notably in Africa.

7. Changing Pleistocene Environments in Africa

8. Changing Pleistocene Environments in Eurasia

9. Key Middle Pleistocene Premodern Human (H. heidelbergensis) Fossils from Africa SiteDates (y.a.)Human Remains Bodo (Ethiopia) Middle Pleistocene (600,000) Incomplete skull, part of braincase Broken Hill (Kabwe) (Zambia) Late Middle Pleistocene; (130,000 or older) Nearly complete cranium, cranial fragments of second individual, miscellaneous postcranial bones

10. Homo heidelbergensis skull from Zambia The Kabwe (Broken Hill) Homo heidelbergensis skull from Zambia. Note the very heavy supraorbital torus.

11. Earliest Evidence of Homo Heidelbergensis in Africa Bodo cranium, the earliest evidence of Homo heidelbergensis in Africa.

12. Key Middle Pleistocene Premodern Human (H. heidelbergensis) Fossils from Europe SiteDates (y.a.)Human Remains Arago (Tautavel, France) 400,000– 300,000; date uncertain Face; parietal perhaps from same person; many cranial fragments; up to 23 individuals represented Atapuerca (Sima de los Huesos, northern Spain) 320,000– 190,000, probably 300,000 Minimum of 28 individuals, including some nearly complete crania

13. Key Middle Pleistocene Premodern Human (H. heidelbergensis) Fossils from Europe SiteDates (y.a.)Human Remains Steinheim (Germany) 300,000– 250,000; date uncertain Nearly complete skull, lacking mandible Swanscombe (England) 300,000– 250,000; date uncertain Occipital and parietals

14. Steinheim Cranium Steinheim cranium, a representative of H. heidelbergensis from Germany.

15. Fossil Discoveries of Middle Pleistocene Premodern Hominids.

17. Key Middle Pleistocene Premodern Human (H. heidelbergensis) Fossils from Asia SiteDates (y.a.)Human Remains Dali (China) Late Middle Pleistocene (230,000– 180,000) Nearly complete skull, lacking mandible. Nearly complete skull Jinniushan (China) Late Middle Pleistocene (200,000) Partial skeleton, including a cranium

18. Crania from China (a) Dali skull and (b) Jinniushan skull, both from China. These two crania are considered by some to be Asian representatives of Homo heidelbergensis.

19. Time line of Middle Pleistocene hominids.

20. Review of Middle Pleistocene Evolution (400,000-125,000 y.a.) Like the erects/sapiens mix in Africa and China, fossils from Europe exhibit traits from both species. Fossils from each continent differ, but the physical differences are not extraordinary. There is a definite increase in brain size and a change in the shape of the skull.

21. Middle Pleistocene Tools African and European archaics invented the Levallois technique for tool making. Acheulian tools are associated with hand axes. Different tool traditions coexist in some areas.

22. The Levallois Technique

23. Culture of Neandertals Neandertals, who lived in the cultural period known as the Middle Paleolithic, are almost always associated with the Mousterian industry. In the early part of the last glacial period, Mousterian culture extended across Europe and North Africa into the former Soviet Union, Israel, Iran, and as far east as Uzbekistan and perhaps even China.

24. Culture of Neandertals Neandertals improved on previous prepared- core techniques by inventing a new variation.  They trimmed a flint nodule around the edges to form a disk-shaped core.  Each time they struck the edge, they produced a flake, continuing this way until the core became too small and was discarded.  They then trimmed the flakes into various forms, such as scrapers, points, and knives.

25. Morphology and Variation in Neandertal Crania

26. Krapina (a) Lateral view showing characteristic Neandertal traits. (b) Three quarters view.

27. Fossil Discoveries of Neandertals

28. Mousterian Tools

29. Settlements People of the Mousterian culture lived in open sites, caves, and rock shelters. Windbreaks of poles and skin were placed at the cave opening for protection against severe weather. Fire was used for cooking, warmth, light, and keeping predators at bay.

30. Excavation of the Tabun Cave, Mt. Carmel, Israel

31. Shanidar 1 Among the individuals buried at Shanidar cave is the skeleton of a one- armed, partially blind, crippled man. He could not have hunted or gathered food on his own. Some believe his survival is proof of Neandertal compassion and humanity

32. Subsistence Remains of animal bones demonstrate that Neandertals were successful hunters. Used close-proximity spears for hunting (spear thrower and bow and arrow weren’t invented until the Upper Paleolithic). Patterns of trauma in Neandertal remains match those of contemporary rodeo performers, indicating close proximity to prey.

33. Symbolic Behavior Prevailing consensus has been that Neandertals were capable of articulate speech. Even if Neandertals did speak, they did not have the same language capabilities of modern Homo sapiens.

34. Burials Neanderthals buried their dead. Their burials included grave goods like animal bones and stone tools. They placed the bodies of their dead in a flexed position.

35. Question The prevailing consensus among researches is that Neandertal: a) was capable of articulate speech. b) probably used grunting sounds to communicate. c) did not have the anatomical features required for speech. d) was unable to speak.

36. Answer: a The prevailing consensus among researches is that Neandertal was capable of articulate speech.

37. Question Regarding deliberate burials: a) Neandertals laid the deceased on their back. b) Neandertals included stone tools. c) Neandertals included a shroud. d) it is seen in Africa before Europe.

38. Answer: b Regarding deliberate burials Neandertals included stone tools.

39. Key Neandertal Fossil Discoveries SiteDates (y.a.)Human Remains Vindija (Croatia) 42,000– 28,000 35 specimens; cranial fragments La Chapelle (France) 50,000Nearly complete male skeleton Shanidar (Iraq) 70,000– 60,000 9 partial skeletons Tabun (Israel) 110,000 date uncertain 2 or 3 individuals, almost complete female skeleton Krapina (Croatia) 125,000 –120,000 Up to 40 individuals, fragmentary

40. Cultural Contrasts: Neandertals and Upper Paleolithic Modern Humans Neandertals Upper Paleolithic Modern Humans Tool Technology Numerous flake tools; few, however, apparently for highly specialized functions; use of bone, antler, or ivory very rare; relatively few tools with more than one or two parts Many more varieties of stone tools; many apparently for specialized functions; frequent use of bone, antler, and ivory; many more tools comprised of two or more component parts

41. Cultural Contrasts: Neandertals and Upper Paleolithic Modern Humans Neandertals Upper Paleolithic Modern Humans Hunting Efficiency and Weapons No long-distance hunting weapons; close-proximity weapons used (more likelihood of injury) Use of spear-thrower and bow and arrow; wider range of social contacts, permitting more organized hunting parties Stone Material Transport Stone materials transported only short distances Stone tool raw materials transported over longer distances, implies wider social networks and trade

42. Cultural Contrasts: Neandertals and Upper Paleolithic Modern Humans Neandertals Upper Paleolithic Modern Humans ArtArtwork uncommon; small; mostly of a personal nature; some items misinterpreted as “art”; others may be intrusive from overlying Upper Paleolithic contexts; cave art absent Artwork much more common, including transportable objects as well as elaborate cave art; well executed,using a variety of materials and techniques; stylistic sophistication

43. Cultural Contrasts: Neandertals and Upper Paleolithic Modern Humans Neandertals Upper Paleolithic Modern Humans Burial Deliberate burial at several sites; graves unelaborated; graves frequently lack artifacts Burials much more complex, frequently including both tools and remains of animals

44. Phylogeny of genus Homo

46. Three Major Evolutionary Transitions 1. Transition from early Homo to H. erectus. Geographically limited to Africa and occurred rapidly. 2. Transition of H. erectus grading into early H. sapiens. Not geographically limited, but occurred slowly and unevenly. 3. Transition from Archaic H. sapiens to anatomically modern H. sapiens.